identifier taxonID type format title accessURI UsageTerms owner agentID CVterm description subtype cb9ba5411cfe20fd17737d82211ae2a8 taxon_achaetodactylus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of Achaetodactylus ceratinae from Ceratina opaca from South Africa. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Achaetodactylus_ceratinae_BMOC_04-0508-286.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2b6ecd46241f3f9eab107832c97bbb1a taxon_achaetodactylus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Achaetodactylus ceratinae seen at the entrances of the axillar acarinaria (arrows) of Ceratina opaca from South Africa.  http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Achaetodactylus_ceratinae_axil_acar_BMOC_04-0508-286.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 661bafcb479f44c440991e729cbb357c taxon_achaetodactylus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Achaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description (unique characters only). Phoretic deutonymph. Free palpi, gnathosomal solenidia and setae absent. Alveoli ve usually weakly developed.
Feeding instars and immobile deutonymph unknown. map_1 taxon_achaetodactylus http://purl.org/dc/dcmitype/StillImage Range map of the Bee-Associated Mite Genus Achaetodactylus http://insectsdataserver.ummz.lsa.umich.edu/bee_mites/index.html?-db=ummz.fm&-format=mapq.js&IDENTITY=Achaetodactylus&-max=200&-find http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database 261193d0fcb6c11b49b2eda76ba564d9 map 78dcf4fd5789b9b3dd98f588f28d17df taxon_achaetodactylus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Achaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Afrotropical region. 2b559df319d87dad4f9d4137f8355522 taxon_achaetodactylus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Achaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Associated with Afrotropical Ceratina spp. Achaetodactylus ceratinae was found in the axillar acarinarium (Fig. 2) (Klimov et al., 2007b) and Achaetodactylus leleupi was found in metasomal acarinarium, suggesting some level of mutualistic relationships between the mites and their hosts. fbf4fa22078cdbb20a79baa939001bba taxon_achaetodactylus_ceratinae http://purl.org/dc/dcmitype/StillImage image/jpeg Achaetodactylus ceratinae, phoretic deutonymph, ex Ceratina opaca, South Africa, BMOC 04-0508-286 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Achaetodactylus_ceratinae_BMOC_04-0508-286.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1443fad0c31d468fe7600c3fbbf19790 taxon_achaetodactylus_ceratinae http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Achaetodactylus ceratinae at the entrances of the axillar acarinaria (arrows) of Ceratina opaca from South Africa. , BMOC 04-0508-286 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Achaetodactylus_ceratinae_axil_acar_BMOC_04-0508-286.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 136870aa7fc1620dc6836a610ff95a13 taxon_achaetodactylus_ceratinae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Achaetodactylus ceratinae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Kenya: Nairobi (type locality), Tanzania, South Africa d3ed4fb62b6876702fea7beb0d5f4756 taxon_achaetodactylus_ceratinae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Achaetodactylus ceratinae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Hirashima) lativentris Friese, 1905 (type host); Ceratina (Hirashima) nigriceps Friese, 1905 (as Ceratina apaca = lapsus for Ceratina opaca) 491812b65277aec64a1eb09ef80148e8 taxon_achaetodactylus_ceratinae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Achaetodactylus ceratinae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the axillar acarinarium of Ceratina nigriceps (Fig. 2) (Klimov et al., 2007b). 9f7423e56e75e9da48aa6e4e448c5798 taxon_achaetodactylus_decellei http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Achaetodactylus decellei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tanzania: Arusha National Park ("de Ngurdoto, Nat. Park") (type locality), Tanzania, Cameroon, Democratic Republic of the Congo 99445d58f49c913d5009527897189bc9 taxon_achaetodactylus_decellei http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Achaetodactylus decellei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina sp. (n° 205 B) (type host); Ceratina spilota Cockerell, 1932 (Cameroon); Ceratina (Propithitis) aereola Vachal, 1903 (Democratic Republic of the Congo); Ceratina excavata Cockerell, 1937 (Tanzania) 0c201b8ef38bf6dda825a6eb041179e8 taxon_achaetodactylus_leleupi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Achaetodactylus leleupi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Kenya: Nairobi (type locality), Tanzania, Cameroon. a10eed07fef435a9d394ab5b3b40b15f taxon_achaetodactylus_leleupi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Achaetodactylus leleupi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina ruwenzorica Cockerell, 1937 (type host); Ceratina diloloensis Cockerell, 1932 (Kenya); Ceratina spilota Cockerell, 1932 (Cameroon); Ceratina (Hirashima) nigriceps Friese, 1905 (Tanzania); Ceratina (Hirashima) lativentris Friese, 1905 (Tanzania) (Fain, 1981b) 9513a575a89fa56e6c06bd33d10275cd taxon_achaetodactylus_leleupi http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Achaetodactylus leleupi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the metasomal acarinarium of Ceratina nigriceps (Fain & Pauly, 2001) f629acb14c5e0a18552f710baeae307f taxon_ascidae http://purl.org/dc/dcmitype/StillImage image/jpeg Proctolaelaps longisetosus, female, ex Bombus fervidus, USA: Ohio, BMOC 02-0706-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Proctolaelaps_longisetosus_BMOC_02-0706-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 22b161ba9dae503218fbe54ed5e58d38 taxon_ascidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Ascidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family is distributed worldwide and includes 34 genera. Ascid mites live in soil, leaf litter, subcortical situations and are often associated with other animals (Halliday et al., 1998). The mites are mostly predators, but also can feed on fungal mycelium; some are parasitic. Several species of the genus Proctolaelaps are obligatory associates of bumblebees. The genera Lasioseiusand Blattisocius were recorded from beehives in North America and the Old World (Basha and Yousef, 2001; Bregetova, 1977; Chinniah and Mohanasundaram, 1995; Crozier, 1989; Haragsim et al., 1978; Malabanan and Corpuz-Raros, 1998). In addition, Melichares agilis Hering, 1838 was found in beehives in Europe (Haragsim et al., 1978), and Antennoseius sp. and Gamasellodes bicolor (Berlese, 1918) were recorded from beehives in Iran (Kamali at al., 2001; Mosaddegh, 1997). 9864bafb9245dc56e3ef1bb25a066006 taxon_blattisocius http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Blattisocius http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Blattisocius comprises at least 10 species, most of which are widespread predators in soil, insect cultures and stored food, and on plants. One species, Blattisocius patagiorum Treat, 1966, is parasitic on adult noctuid moths (Halliday et al., 1998). Blattisocius tarsalis (Berlese, 1918) was found in honeybee hives (Apis mellifera) in Canada (Crozier, 1989). This and three others species, Blattisocius dentriticus (Berlese, 1918), Blattisocius apisassociae Chinniah et Mohanasundaram, 1995, and Blattisocius apis Basha et Yousef, 2001 were recorded from the same habitat in the Old World (Banaszak, 1980; Basha and Yousef, 2001; Chinniah and Mohanasundaram, 1995; Haragsim et al., 1978). In addition, Lasioseius sp. was collected in the Philippines from Apis cerana and Apis mellifera (Malabanan and Corpuz-Raros, 1998). 7ab22b059f607fd045812bcf484a60ed taxon_blattisocius_tarsalis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Blattisocius tarsalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Nearctic (including the United States and Canada), Palaearctic (Italy is the type locality), Oriental, and Australian regions (Berlese, 1918; Bhattacharyya and Sanyal, 2002; Karg, 1971) 73064cf619c4d6bdbddf7f914f12fae8 taxon_blattisocius_tarsalis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Blattisocius tarsalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species commonly occurs is synanthropic habitats (hay, stored products). In Western Australia, it was found on larvae and pupae of the greater wax moth Galleria mellonella (Linnaeus, 1758) in beehive (Halliday et al., 1998); Crozier (1989) reports it from honey bee hives in Canada. In North America, it was also recorded from cultures of various stored product insects: Sitotroga cerealella(Olivier, 1789) (Lepidoptera: Gelechiidae), Tineola biselliella (Hummel, 1823) (Lepidoptera: Tineidae), Ephestia elutella (Hübner, 1796) (Lepidoptera: Pyralidae); Tribolium confusum Jacquelin du Val, 1863, Tribolium castaneum (Herbst, 1797) (Coleoptera: Tenebrionidae), Attagenus piceus Latreille, 1804 (Coleoptera: Dermestidae). It was shown that Blattisocius tarsalis could control the size of the Mediterranean Flour Moth (Ephestia kuehniella Zeller, 1879) populations in specific conditions (depth of grain); the mites consumed the insect eggs and were phoretic upon the adult moths. Blattisocius tarsalis may be a parasitic species, Treat (1975) noticed attacks of the female mites on full grown larvae of the moth Idia aemula (Hübner, 1813) (Lepidoptera: Noctuidae). He also lists this species of moth, as well as Apamea devastator Brace, 1819 (Noctuidae) and Aglossa costiferalis (Walker, 1866) (Pyralidae) as phoretic hosts for Blattisocius tarsalis in the United States. The mite life span from egg to egg is approximately ten days.
7be9caa21ee10173278bd8581b62fad5 taxon_centriacarus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Centriacarus guahibo (holotype) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_guahibo_BMOC_04-0508-239.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3f145e87d95b803565b65b7ad0d985fe taxon_centriacarus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Centriacarus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description (unique character states underlined). Phoretic deutonymph. Gnathosomal solenidion, free palpi and their setae present. Alveoli ve dorsal, distinctly anterior to se. Prodorsal shield striation longitudinal anteriorly and transverse posteriorly. Prodorsal shield present. Posterior edge of prodorsal shield shorter than its lateral edges. Setae se situated on soft cuticle. Setae c2 situated on same transverse level as c1. Setae e2 situated on hysterosomal shield. Setae 1a and 3a touching posterior borders of respective coxal fields and filiform. Cupules ia situated on hysterosomal shield. Cupules im distinctly posterior to acetabula III, situated off line between d2 and e2. Cupules ip anterior to setae f2. Cupules ih situated on sides of attachment organ. Posterior part of posterior apodemes of coxal fields II not displaced posteriorly to anterior apodemes III. Coxal fields III closed. Coxal fields IV open. Transverse medial extension of posterior apodemes IV well-developed. Anterior extension of posterior apodemes IV present, connecting with anterior apodeme III. Ventral longitudinal sclerites of progenital chamber conspicuous at posterior part. Ventral longitudinal sclerites of progenital chamber conspicuous at anterior part. Posterior and lateral cuticular suckers (Fig. 2) present. Anterior cuticular suckers present (Fig. 2). Bases of anterior cuticular suckers inserted on separate apodeme (may touch or overlap posterio-lateral sclerotized border of the attachment organ) (Fig. 2). Apodemes of ps1 separated. Setae wa I-II submedial, f I-II apical, near tarsal apices. Solenidion ω2 present. Empodial claws I-III not twisted. Dorsal cuticular folds of ambulacra I-III absent (Fig. 3). Condylophores of tarsi I-III weakly developed, almost symmetrical. Supporting sclerites of condylophores (latero-apical sclerites of tarsus) indistinct from tarsus, not connected by dorsal bridge (Fig. 4). Disto-dorsal lobe of distal part of caruncle absent. Dorsal condylar plate of femur-tibia joint broad. Tarsi I-II with 7 setae (p and q present). Tarsal setae ra and la I-II foliate. Genual seta cG I longer or only slightly shorter than genu I and modified. Genual setae cG I longer than cG II. Tarsal setae q III present. Tarsal setae w, r, and p III present. Tarsal seta s III foliate. Sigma III absent, represented by alveolus. Tarsus IV with 8 setae (s, p, q present). Tarsal setae e, f IV foliate or slightly lanceolate. Tarsal setae w IV longer than leg IV. Tibial setae kT IV present. Solenidion φ IV absent, represented by alveolus.
Feeding instars and immobile deutonymph unknown.
Notes. Feeding instars of the Sennertia vaga complex have been collected on the same hosts, sometimes together with deutonymphs of Centriacarus. They have poorer leg chaetotaxy and belong to an early derivative lineage that probably does not form deutonymphs. 6013c3ebae3d496182fdae59e4456cd2 taxon_centriacarus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Centriacarus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Neotropical region. 9c695db13addb81f5df8c47f21f8f6ad taxon_centriacarus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Centriacarus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Associated with Centris (Heterocentris). 666d9ccea06f84b6145ced1998f15215 taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Centriacarus guahibo (holotype, BMOC 04-0508-239) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_guahibo_BMOC_04-0508-239.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b54bf7d776f4a83683cc8916d5361525 taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Centriacarus guahibo on the bee, Centris in Venezuela (museum specimen), BMOC 91-0402-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_guahibo_on_Centris_BMOC_04-0508-239.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 49dc7def9ea6dfce412c9f30c3d2c7bc taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Centriacarus guahibo on the bee, Centris in Venezuela (museum specimen), BMOC 91-0402-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_guahibo_on_Centris_BMOC_04-0508-239b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7f76b739398ec1802c62ea79b80e73b3 taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Centriacarus guahibo http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Setae c1 usually reaching or extending beyond bases of d1, c1/c1-d1 1.1 0.7-1.3 (1.0±0.17). Setae 3a and 1a longer than 4b, distinctly shorter than c3. Ratio 3a/c3 1.0-2.0 (1.4±0.19), 1a/c3 1.5-2.1 (1.7±0.16). Setae 4b nearly equal to c3, 1a/c3 0.7-1.2 (1.0±0.12). aec7e508867d4e237da4b43c039e9ff9 taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Centriacarus guahibo http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Venezuela: Territorio Amazonas. 838d68642c4e9e56721f87289d8090b6 taxon_centriacarus_guahibo http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Centriacarus guahibo http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Centris sp. a6e9b3a2370fc03da547f7d1bb51e74c taxon_centriacarus_turbator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of the mite Centriacarus turbator (holotype), ex Centris vittata, Brazil. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_turbator_ventr_BMOC-95-0422_026.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 69653271b7233bd0886f19027fa20671 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of the mite Centriacarus turbator (dorsal view), ex Centris vittata, Brazil http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_turbator_BMOC_95-0422-026.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 09cb1b768dd4bdd0f24a2672e89d4fe6 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/StillImage image/jpeg Centris sp. with phoretic deutonymphs of Centriacarus turbator, Mexico (BMOC 04-0508-237) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_turbator_BMOC-04-0508-237.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 52005eda3075eda83917a9bdb439fbf3 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Centriacarus turbator on propodeum of Centris sp., Mexico (BMOC 04-0508-237) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_turbator_BMOC-04-0508-237c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 64442c3eb7c49bee86aecd66f224eb58 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Centriacarus turbator on 1st metasomal tergite of Centris sp., Mexico (BMOC 04-0508-237) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Centriacarus_turbator_BMOC-04-0508-237b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3ada7214ed6b3d8720838083f8bdf0ea taxon_centriacarus_turbator http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Centriacarus turbator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph (Fig. 1). Setae c1 usually not reaching bases of d1, c1/c1-d1 0.7-1.0 (0.8±0.09). Setae 3a and 4b distinctly shorter than c3. Ratio 3a/c3 0.4-0.7 (0.5±0.07), 4b/c3 0.4-0.8 (0.6±0.10). Setae 1a nearly equal to c3, 1a/c3 0.8-1.2 (1.0±0.13). a9ef2ed8d3103dc297dbca30f802d5b6 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Centriacarus turbator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Mato Grosso do Sul (type locality), Peru, Colombia, Panama, Mexico: Jalisco. 3e974ff769a6fa9f5c005ecfcbba7db2 taxon_centriacarus_turbator http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Centriacarus turbator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Centris (Heterocentris) vittata (type host), Centris sp. (Mexico) 83d430478853f18565a30f04b5fc05de taxon_chaetodactylidae http://purl.org/dc/dcmitype/StillImage image/jpeg Mite Chaetodactylus krombeini (Chaetodactylidae). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_krombeini.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 182a076a80c515bcb1f3f2da99422ee6 taxon_chaetodactylidae http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Family Chaetodactylidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The family-group name is a junior homonym of Chaetodactylini Tschitscherin, 1903 (Coleoptera) (OConnor, 1993). 4fbc8774de5c3ac01d04b41c9ed60301 taxon_chaetodactylidae http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Family Chaetodactylidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription The supracoxal sclerites are enlarged and modified. External vertical setae ve are absent or reduced to alveoli in all instars. Tarsal setae aa I, u and v I-IV are absent from all instars. Solenidion ω3 is shifted to the posterior part of tarsus I in tritonymphs and females (in males it is on the anterior side). In non-deutonymphs, the anterior oblique ridge of the gnathosoma is well-developed, starting near the posterior transverse ridge and extending anteriorly, meeting the internal wall of the palpcoxae. In heteromorphic deutonymphs, supracoxal setae scx are vestigial, with rounded or blunt tips; setae e and ba I-II are absent. 6c17a5c723761c857f8990166603148e taxon_chaetodactylidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Chaetodactylidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes 5 genera more than 112 species and subspecies distributed worldwide (show map), except for Antarctica (Klimov and OConnor 2008). Chaetodactylids are obligate associates of long-tongued bees and has developed striking morphological, developmental, and biological adaptations to its hosts. The mites live in nests of bees as mutualists (feeding on nest waste), parasitoids (killing the bee egg or larvae), commensals or cleptoparasites (feeding on provisioned pollen) (Krombein 1962, Qu et al. 2002, Roubik 1987). Some parasitoids and cleptoparasites cause substantial damage to managed colonies of mason bees (Osmia) used as commercial pollinators (Bosch 1992, Bosch and Kemp 2001, Fain 1966, Kurosa 1987, Park et al. 2009).
Chaetodactylids disperse as heteromorphic deutonymphs on the adult insects, and the life cycles of the mites and their hosts are usually synchronized. In the Sennertia vaga species group, the deutonymph is probably lost and the mites disperse as feeding instars on adults of large carpenter bees. This group is unusual among astigmatid mites in that reproduction and feeding may occur during dispersal.
At least some species of the genus Chaetodactylus are able to produce morphologically regressive, non-phoretic, heteromorphic deutonymphs. These remain in the nest cavity to infest a new generation of cells when the cavity is re-used.
Most chaetodactylids are oligoxenous, utilizing several closely related hosts and monophyletic groups of mites are often restricted to monophyletic groups of bees, suggesting that they may share common coevolutionary histories. However, historical dispersal of mites to new hosts, most probably, were governed by ecological factors (Klimov et al. 2007a), such as the bee nest architecture. Several bee species of the genera Tetrapedia, Ceratina and Xylocopa have even developed special pouches for mite transfer (acarinaria) (Klimov et al. 2007b), indicating possible mutualistic relationships.
Central and South America, the probable center of origin of the family, are characterized by an exceptional concentration of endemic lineages of mites and their hosts, unfortunately only a few mite species have been described from these regions (Klimov and OConnor 2007). 066a5b7291548586031c2562d52076df taxon_chaetodactylus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus krombeini dispersing on the orchard mason bee, Osmia lignaria, Michigan http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_krombeini_BMOC_06-0508-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ce07563b934a6ecbd3d8a30fd785ecaa taxon_chaetodactylus http://purl.org/dc/dcmitype/StillImage image/jpeg Immobile heteromorphic deutonymph of Chaetodactylus micheneri. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_micheneri_immob.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 30d1789904d5dbf1ffe436559fdc375a taxon_chaetodactylus http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Genus Chaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription (unique characters only). Phoretic deutonymph. Gnathosomal solenidion present and setae on free palpi absent and free palpi present (unique combination of character states). Immobile deutonymph present. Adults. Cupules im ventral. Male. Progenital sclerites completely fused forming large unpaired sclerite (Fig. 3). Lateral processes (horns) of dorsal supporting sclerite with secondary processes (Fig. 4). Distinct anterio-dorsal protuberance on tarsi I-IV present (Fig. 5). Heteromorphic males absent. Larva. Claparède's organ shaft more or less cylindrical, constricted apically; dome spherical. 291b2afd3f72bc19d399bf28cc3cb0b8 taxon_chaetodactylus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Chaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Chaetodactylus is associated with megachilid bees on a worldwide basis (except Antarctica), while species associated with apid bees are only found in the Neotropical region. Chaetodactylus is associated with Hoplitis species in the Nearctic region, but not in the Old World where this host bee genus also occurs. Similarly, no records are known for Chaetodactylus associated with North American Chelostoma and Megachile, while the mites do occur on these hosts in the western Palaearctic. The close similarity of species of the osmiae-lineage and Ch. anthidii associated with Rhodanthidium sticticum suggests a recent host shift from Osmia and subsequent vicariance in the Palaearctic region. No Chaetodactylus are positively known from New World Anthidiini, although we were able to find one on Anthidium spp. in Chile. The distribution patterns of Ch. krombeini + Ch. claviger and Ch. claudus + Ch. osmiae, sister species living in the Nearctic and southwestern Palaearctic regions, respectively, indicate their recent vicariance. The ludwigi-dalyi lineage has a broad distribution in the Australian, Oriental, Afrotropical, Neotropical, and south Palaearctic regions, which may imply intercontinental dispersal events or an ancient Gondwanan origin of this early derivative lineage. f09050eab3a43693027cca14604a7004 taxon_chaetodactylus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Chaetodactylus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Species of this genus are associated with Megachilidae, tribes Lithurgini (Lithurgus, Trichothurgus, Microthurge), Osmiini (Osmia (Fig. 1), Hoplitis, Chelostoma), Anthidiini (Rhodanthidium, Anthidium), and Megachilini (Megachile), and with Apidae, tribes Emphorini (Melitoma, Diadasia, Ptilothrix, Ancyloscelis), and Tapinotaspidini (Chalepogenus). Most species occur on Lithurgus and Osmia, while only one species is associated with each of the remaining host genera.

Fig. 2. Immobile heteromorphic deutonymph of Chaetodactylus micheneri. Click here to enlargeThe mites usually kill young bee larvae and feed on provisioned pollen and nectar. In nests with partitions (Osmia), bees that develop in the innermost cells chew their way out of the nest, and phoretic deutonymphs from the opened cells may attach to them. The mites in the innermost cell would possibly die because of their inability to break through the partition. In nests without partitions (Lithurgus), some young bees possibly complete development and transform to adults that disperse the mites.
The presence of the inert non-phoretic deutonymph along with the phoretic deutonymph is the most conspicuous feature in the life-cycle of this genus. This is a highly regressive, cyst-like morph with legs and most setae greatly reduced (Fig. 2). It is capable of surviving in old bee nests and infesting new hosts that reuse these nests or nest material. Inert deutonymphs are very important for the mite survival when mites are trapped in innermost cells of an infested nest or all bee larvae are killed and therefore cannot transfer mites to a new nest as adults.
Biology has been studied for Chaetodactylus osmiae (Chmielewski 1993, Fain 1966, Lith 1957, Popovici-Baznosanu 1913), Ch. birulai (Lith 1957), and Ch. krombeini (Krombein 1962, 1967). e5236c88d957c4f24b639d4762a1bccc taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus abditus, phoretic deutonymph (holotype), ex Lithurgus planifrons, Arizona, BMOC 96-0510-012 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_abditus_holotype_BMOC_96-0510-012.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e cc98c38624e39d3fcaf1a26238f2c56e taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus abditus, phoretic deutonymph (holotype) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_abditusB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b27519f0d71feeb12e59d847c3ced989 taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus abditus on Lithurgus echinocacti pollinating barrel cactus (Ferocactus), Arizona http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_abditusB_Lithurgus_Lithurgus_echinocacti.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e fb6ac6042cc577a9aabb416f92d52260 taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus abditus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona (type locality), Mexico: Socorro Island. 912a668f722a08e0970a4080dd3358d0 taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus abditus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations ; Lithurgus (Lithurgopsis) planifrons Friese, 1908 (type host); Lithurgus (Lithurgopsis) echinocacti Cockerell, 1898;
5a542dde07c951ea5733edf636080125 taxon_chaetodactylus_abditus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus abditus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology In Arizona is found phoretic on bee species of the genus Lithurgus flying in the fall. This contrasts to the sympatric species Chaetodactylus lithurgi associated with Lithurgus species which are active during the spring. 01221f856245ef2ddae44c8c39e10810 taxon_chaetodactylus_anthidii http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus anthidii, phoretic deutonymph, ex Rhodanthidium sticticum, France, BMOC 79-0329-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_anthidii_BMOC_79-0329-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2b18ba3d7d0f7281ce1ee7bd8418e73e taxon_chaetodactylus_anthidii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus anthidii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tunisia (type locality), France (Oudemans, 1911a; Klimov & OConnor, 2008) 5c65d64b6b8a1cbce32a4354164bcaf2 taxon_chaetodactylus_anthidii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus anthidii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Rhodanthidium sticticum (Fabricius, 1787) (as Anthidium sticticum) aeac0882356e551a3f64481ce9c803c3 taxon_chaetodactylus_antillarum http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus antillarum, phoretic deutonymph (holotype), ex Lithurgus antilleorum, Jamaica, BMOC 96-0916-190 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_antillarum_BMOC_96-0916-190.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f24e2a6d553875d51d852c371b0006ec taxon_chaetodactylus_antillarum http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus antillarum http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_antillarumB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 68b5cb038ac1ade5b9c98d338236bde3 taxon_chaetodactylus_antillarum http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus antillarum on a museum specimen of Lithurgus antilleorum, Jamaica, BMOC 96-0916-190 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_antillarumB-Lithurgus_antilleorum_BMOC_96-0916-190.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 353b7d38c073562a5f1e875cdeff42d9 taxon_chaetodactylus_antillarum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus antillarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Dominican Republic: Pedernales; Jamaica: Parish of Saint Catherine (type locality). 755ea561b91d7294698c0981836a7308 taxon_chaetodactylus_antillarum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus antillarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgopsis) antilleorum antilleorum (Megachilidae). d0a36757697b5b84b73f3de60e734723 taxon_chaetodactylus_azteca http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus azteca, phoretic deutonymph (holotype), ex Osmia azteca, Mexico, BMOC 96-0510-139 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_azteca_BMOC_96-0510-139.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 890c6a12a1cc25d8f16bde2b86efc84a taxon_chaetodactylus_azteca http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus azteca http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_aztecaB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c577c6a9a6f354faed4c66b0a8e1f3fd taxon_chaetodactylus_azteca http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus azteca on a museum specimen of the bee Osmia azteca from Mexico http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_aztecaB-BMOC-96-0510-140.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a15a4764d1f53906d35dc321a1fcfeca taxon_chaetodactylus_azteca http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus azteca http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Chiapas, Hidalgo (type locality). af00069f3fcb9cfb32345db11d7c3046 taxon_chaetodactylus_azteca http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus azteca http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Diceratosmia) azteca. 07890d00d03a2516a772c84ea62fb5b9 taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus birulai, phoretic deutonymph (lectotype), ex Chelostoma florisomne, Russia, BMOC 11-0618-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_birulai_lectotype_dors_BMOC_11-0618-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 143a408c756f1c8c0bcff6646ef3d6c6 taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus birulai syntype HDN, ventral view http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_birulai_lectotype_ventr_BMOC_11-0618-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9418a160d91d0ab2a77b288d4e77a776 taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus birulai syntype http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_birulai_lectotype_slide_BMOC_11-0618-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 5273b8ad6383dc60d3e70db4219ca157 taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus birulai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Russia: Kirovskaya Oblast' ("Lavy") (type locality) (Zachvatkin, 1941); Netherlands (Lith, 1957); Czech Republic (Samšiňák, 1973), Austria (our data) 0b733e127f4b9b8abad4126fb29fe0e4 taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus birulai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Chelostoma florisomne (Linnaeus, 1758) (as Chelostoma florisomnis) 1755addbf5586a8b16ecb8bf48cc96db taxon_chaetodactylus_birulai http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus birulai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology May kill bee larvae in the nest (Lith, 1957). 9265e9c470ecf71daeccd83ad7261743 taxon_chaetodactylus_chrysidis http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus chrysidis, phoretic deutonymph (holotype), ex Osmia aurulenta, Germany, BMOC 95-0315-012 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_chrysidis_BMOC_95-0315-012.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1ab23ffd25d3849c409293a960fa8947 taxon_chaetodactylus_chrysidis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus chrysidis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Belgium: "Treignes-Saumières" (type locality) (Fain & Baugnée, 1996), Germany (Klimov & OConnor, 2008) a49eab741a0ec3d787c436a756c1f67e taxon_chaetodactylus_chrysidis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus chrysidis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Chrysura trimaculata (Förster) (Hymenoptera, Chrysididae) (type host), parasitoid of Osmia aurulenta (Panzer, 1799) and O. bicolor (Schrank, 1781) 61dea691fd6a6603de472878cdc7cc3f taxon_chaetodactylus_claudus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus claudus, phoretic deutonymph (holotype), ex Osmia ribifloris, Utah, BMOC 96-0510-092 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_claudus_BMOC_96-0510-092.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 6f388318f0bca52f81a5f87b41bb3581 taxon_chaetodactylus_claudus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus claudus, ventral and dorsal views, phoretic deutonymph (holotype) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_claudusB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 5a4722d7dcf0ea0ba651c88bd982ba72 taxon_chaetodactylus_claudus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus claudus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Texas, Utah. 4414fe0bd2806a30763bf58176ae9618 taxon_chaetodactylus_claudus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus claudus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Osmia) ribifloris (Megachilidae). 00877078c4801af3dc4e7dd6042d33d6 taxon_chaetodactylus_claviger http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus claviger, phoretic deutonymph, ex Osmia tricornis, Italy, BMOC 03-0523-006 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_claviger_BMOC_03-0523-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 09d3de615e86a26f213530772ace919e taxon_chaetodactylus_claviger http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus claviger http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Italy: Promontorio del Gargano ("Monte Gargano, Apulië") (Oudemans, 1924); France (Donnadieu, 1868); Egypt (Abou Senna, 1997) 73eccc29fa39fdb0ea06faf36a065b00 taxon_chaetodactylus_claviger http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus claviger http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia tricornis Latreille, 1811; Apis mellifera (Egypt) (accidental record) 278d6412cf95597fd4b81473c0a9f8cd taxon_chaetodactylus_dalyi_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus dalyi ( http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution South Africa: "Natal" (type locality), Limpopo ("Transvaal"), KwaZulu-Natal (Saint Lucia "Sta Lucia, Natal"), Zimbabwe (Fain, 1974a) or Mozambique (Fain, 1981b) was inconsistently indicated for the same paratype specimen ex Ceratina sp. (bee no 285); Mozambique (Klimov & OConnor , 2008). f9e40931201869c6a80b4c0ed9b62bdf taxon_chaetodactylus_dalyi_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus dalyi ( http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Pithitis) inermis Friese, 1905 (as Ceratina turneri) (type host, South Africa) (Fain, 1974a); Ceratina sp. (Zimbabwe or Mozambique) (Fain, 1974a; Fain, 1981b); Lithurgus pullatus Vachal, 1903 (Mozambique) (Klimov & OConnor , 2008); Lithurgus aethiops (not found in ITIS) (Mozambique) (Klimov & OConnor , 2008) 7447ec6db4bd342d19f9f884a73acb85 taxon_chaetodactylus_dementjevi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus dementjevi, phoretic deutonymph, ex Chalicodoma bombycina, Russia: Leningradskaya Oblast', BMOC 96-0510-196 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_dementjevi_BMOC_96-0510-196.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e244c67fff81201741e7261164f94b51 taxon_chaetodactylus_dementjevi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus dementjevi, phoretic deutonymph, ex Chalicodoma bombycina, Russia: Leningradskaya Oblast http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_dementjevi_BMOC_96-0510-196b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8e15a17fb4c33ebed469e40ad5b7b231 taxon_chaetodactylus_dementjevi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus azteca http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_dementjevi_syntype_slide_BMOC_11-0618-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 330b27857c26ffaafb6e275223e9a119 taxon_chaetodactylus_dementjevi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus dementjevi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Russia: Leningradskaya Oblast', Yaroslavskaya Oblast'; Kazakhstan: Shyghys Qazaqstan (Ust'-Kamenogorsk); Uzbekistan: Toshkent (Gora Aktash) (Zachvatkin, 1941 - Syntype localities) 87b190561e508e808cf39e316b65a9ad taxon_chaetodactylus_dementjevi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus dementjevi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Megachile bombycina Radoszkowski, 1874 ; Megachile ligniseca (Kirby, 1802) ; Paravespula germanica (Linnaeus) (Vespidae) (accidental host) (Uzbekistan) e9c9c571964b5aea23b18583c98dc050 taxon_chaetodactylus_furunculus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus furunculus, phoretic deutonymph, ex Lithurgus listrotus, California, BMOC 04-1122-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_furunculus_holotype_BMOC_04-1122-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e426c0581bffa26fcf2c026e7e9e75f6 taxon_chaetodactylus_furunculus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus furunculus http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_furunculusB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f4ea683adb76feaa0131bebfd0e03ea3 taxon_chaetodactylus_furunculus http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus furunculus on a museum specimen of Lithurgus listrotus, California http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_furunculusB_BMOC-04-1122-002a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1db3c6cb05a06c124d6318e41244b44b taxon_chaetodactylus_furunculus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus furunculus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: California. ae6df62a0c5a3555e4b7d525925b70d9 taxon_chaetodactylus_furunculus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus furunculus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgopsis) listrotus (Megachilidae). 1339f624abd6453acda62e720a052537 taxon_chaetodactylus_gibbosi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus gibbosi, phoretic deutonymph (holotype), ex Lithurgus gibbosus, Florida, BMOC 02-1205-006 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_gibbosi_holotype_BMOC_02-1205-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bb2ee2defb93dabc882118e9d29fe88f taxon_chaetodactylus_gibbosi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus gibbosi, phoretic deutonymph, holotype, dorsal and ventral, BMOC 02-1205-006 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_gibbosiB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 806577db2a40857c55bad5670156fd09 taxon_chaetodactylus_gibbosi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus gibbosi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Florida (type locality), Georgia. f72a77a9f35da126d85d76c7a7758b54 taxon_chaetodactylus_gibbosi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus gibbosi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgopsis) gibbosus Smith, 1853 d2d77ca56a91ba3f129034602e1b3e5e taxon_chaetodactylus_hirashimai http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus hirashimai, phoretic deutonymph, ex Osmia excavata, Japan, BMOC 04-0508-004 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_hirashimai_BMOC_04-0508-004.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a6a70ce89fcb2d2927a17700d68a6cda taxon_chaetodactylus_hirashimai http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus hirashimai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan: Honshu (Nara Pref., Nara-shi, Furuichi-chô) (type locality), Kyushu d78076e01cc853a85fd648043967d998 taxon_chaetodactylus_hirashimai http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus hirashimai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Osmia) excavata Alfken, 1903 (type host); Osmia (Osmia) cornifrons (Radoszkowski, 1887) ; Osmia (Osmia) pedicornis Cockerell, 1919 ; Osmia (Helicosmia) jacoti Cockerell, 1929 (as Osmia (Helicosmia) imaii) 42cf14115e33aff55421afc551f86974 taxon_chaetodactylus_hirashimai http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus hirashimai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Pest of Osmia excavata used as pollinator in orchards in Japan. If the number of mites in an infested cell is more than 50, then can kill the bee egg or developing larva (up to the early third instar) (Qu et al., 2002). c327e7479204b9c9d06dba5d72fcd8c8 taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus hopliti, phoretic deutonymph (holotype), ex Hoplitis producta, New York, BMOC 84-0409-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_hopliti_holotype_BMOC_84-0409-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c2bfce1b26970ea8e9044de59714454a taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus hopliti, ventral and dorsal view (holotype) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_hoplitiB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3fac9c45b5ec31d81a8c54126e4864d6 taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Chaetodactylus hopliti, on a museum specimen of the bee Hoplitis producta (Megachilidae), BMOC 03-1106-037 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_hoplitiB_BMOC-03-1106-037.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c632302c16a4bcd978ecef4119120019 taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus hopliti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The association of Ch. hopliti and the non-Hoplitis hosts (see above) is probably accidental. Because these insects construct their nests in the same habitat as Hoplitis, it seems possible that deutonymphs of Ch. hopliti may accidentally attach to them. 7686c5009389f23e61be714123a4c204 taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus hopliti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Idaho, Maine, Michigan, New York (type locality), North Carolina, Ohio, Oregon, Tennessee, Wisconsin; Canada: Quebec. 0b52706ba48a8c0a3107306d1764c5ec taxon_chaetodactylus_hopliti http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus hopliti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Hoplitis (Alcidamea) producta (type host); Hoplitis (Alcidamea) pilosifrons; Hoplitis (Monumetha) spoliata; Megachile (Megachile) relativa; Osmia (Melanosmia) brevis; Osmia (Centrosmia) bucephala; Osmia (Chenosmia) inermis (Megachilidae); Isodontia mexicana (Hymenoptera: Sphecidae) 57da67dec0dd30982c771a18259c6a13 taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus kouboy, phoretic deutonymph, holotype, ex Lithurgus apicalis, New Mexico, BMOC 95-0323-020 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_kouboy_holotype_BMOC_95-0323-020.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 03f2592e1f23648fc7334e882d5a764e taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus kouboy http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_kouboy.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 68223fadcc20dfd38eebdc852836f63c taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the Chaetodactylus kouboy and Chaetodactylus lithurgi on a museum specimen of Lithurgus apicalis, New Mexico http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Lithurgus_apicalis_BMOC-_95-0323-020b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2b7c644f9365a21f68f786df512ca9a5 taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus kouboy http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Chaetodactylus kouboy was found only once, on a host also harboring C. lithurgi. This suggests that the host association may be accidental. 713f9451432f8410935bfc42739e9333 taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus kouboy http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: New Mexico. 0cc91a1bde4f97754e8da268046d9da6 taxon_chaetodactylus_kouboy http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus kouboy http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgopsis) apicalis (Megachilidae). 23e21620f554f44af2c75accb704f8a9 taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus krombeini dispersing on the orchard mason bee, Osmia lignaria, Michigan. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_krombeini_BMOC_06-0508-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 997169a133efeb456bc77ae5af2ae67e taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/StillImage image/jpeg Mite Chaetodactylus krombeini (Chaetodactylidae) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_krombeini.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 389eacc4512efd199297ffc8683d52a3 taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus krombeini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Specimens from Osmia pumila (8 HDNs, 4 females - USA: New York, mixed sample from 5 localities from Suffolk (4) and Nassau Co. (1), Osmia pumila nest cell V-VI 1998, K. Goodell UMMZ BMOC 98-1110-001) probably belong to a closely related species. In three well mounted females, the dorsal opisthosoma has a pattern of large tubercles (1.9-2.5 in diameter) (versus conical or subconical mammillae (1.5-2.0) in Ch. krombeini) and the outer ridge of the supracoxal sclerite is shorter than tibia II (longer in Ch. krombeini). Deutonymphs from O. pumila do not have any distinct differences from those of Ch. krombeini, but all have slightly thinner s III and longer e1. Since mites from O. pumila originated from mixed samples with small sample size, and all characters of phoretic deutonymphs overlap with Ch. krombeini, we refrain from specifically determining these specimens at this time. 4732c545f9b03990145a53b0cab1676b taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus krombeini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, California, Idaho, Maryland (type locality), Michigan, Nevada, New York, Ohio, Oregon, Utah, Washington. 4b6f519686e9c2665d2adf694b33af46 taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus krombeini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Osmia) lignaria Say, 1837 (type host); Osmia (Osmia) lignaria propinqua Cresson, 1864; Osmia (Osmia) ribifloris Cockerell, 1900; Osmia (Osmia) ribifloris biedermannii Michener, 1936; Osmia (Acanthosmioides) nigrifrons Cresson, 1878; Osmia (Centrosmia) bucephala Cresson, 1864; Osmia (Cephalosmia) californica Cresson, 1864; Osmia (Cephalosmia) montana Cresson, 1864; Osmia (Helicosmia) chalybea Smith, 1853; Osmia (Melanosmia) grindeliae Cockerell, 1910; Osmia (Melanosmia) simillima Smith, 1853 7390d56448c3a2656cfcb5cf2b72a7e8 taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus krombeini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Chaetodactylus krombeini is primarily associated with the megachilid bee Osmia lignaria. Life cycles of both bee and mite were studied by Krombein (1962). He observed deutonymphs disposed in a random fashion over the bee's body, although the majority were attached to setae on the propodeum or anterior part of the metasoma. One or more of these deutonymphs were observed to crawl off the body of the female bee while she provisioned the cell with pollen and nectar. Presumably they then transformed into tritonymphs. Adult mites of both sexes were present in infested cells 3 to 4 days after the cell was provisioned. In an attempt to explain this phenomenon, Krombein speculates that the phoretic deutonymphs transform into tritonymphs and females, each of which lays a single egg that develops very rapidly into an adult male. This male mates with its mother, or with another female that may be in the same cell, and the female then proceeds to lay fertilized eggs.
Male bees were more commonly infested with phoretic deutonymphs, and also had more mites per bee. This phenomenon is possibly a consequence of the skewed sex ratio and of the prior emergence of male bees in the spring.
Chaetodactylus krombeini may attack and kill the egg or young larva in a newly provisioned cell or in newly infested cells. Occasionally the mites do not kill the host but feed on the provisioned pollen, and the young bee larva is nutritionally deprived (Bosch and Kemp 2001). In this case a smaller than normal adult bee may develop, along with some mites. Ordinarily, the mites are unable to gain access to uninfested cells once the infested cell is capped, and they are confined to the original cell until the partition is broken down the following spring by emergence of an adult bee from one of the earlier constructed cells. Mites frequently move into adjacent cells when partitions break during nest manipulation in managed colonies of Osmia lignaria (Bosch and Kemp 2001).
After killing and feeding on the bee egg or young larva, the female mite deposits her eggs principally on the cell walls beyond the pollen-nectar mass. The eggs hatch in 4 to 5 days into larvae. The larvae feed on nectar from the pollen-nectar mass and transform into protonymphs, which also continue to feed on the nectar. There is some doubt as to what happens next, but it seems probable that the protonymphs occurring early in the season transform directly into tritonymphs, bypassing the deutonymphal stage completely. However, Chaetodactylus deutonymphs were never found in nests early in the spring. The tritonymphs transform into adults, which in turn repeat the cycle, within the infested cell until all the provisioned food has been consumed. The number of generations and duration of breeding is dependent on the volume of the pollen-nectar mass.
Formation of the immobile deutonymph in Chaetodactylus may be due to insufficient food or to decreased humidity caused by the use of all of the nectar, or to a combination of both factors. It is not possible to state what factors determine whether immobile and/or phoretic deutonymphs will develop in an individual nest. Formation of the latter was never observed in the laboratory. In an undisturbed nest, both forms of deutonymphs are probably confined to the originally infested cells because of the presumed inability of the mites to break through the mud partitions separating the cells.
Very early in the spring the phoretic deutonymph presumably attaches to an adult bee as the latter chews its way through the mud seal capping its cell. In an undisturbed nest the mites in the innermost cell or cells would possibly die in situ because of their inability to mount an adult bee. The mites would need to infest some cells in the middle or near the outer end of the nest, so that bees would develop in the innermost cells and provide the necessary vehicle for migration of the mites as the bees chewed their way out of the nest. Mites trapped in the innermost cell might be released by a female bee chewing through the closing partition during her efforts to clean out the debris from an old nest for re-use.
The role of the immobile deutonymph in initiating a new infestation requires additional investigation. Some of the immobile deutonymphs transformed to tritonymphs several days after the bees left the nest in the spring. Theoretically, it would be possible for the immobile deutonymphs to remain in that stage in an old nest for some length of time. If this nest was then re-used by another bee, the capping of cells by that bee would increase the humidity to the point where the heteromorphic deutonymphs could transform into tritonymphs, which would then infest the cells provisioned by that bee.
Obviously, the presence of both immobile and phoretic deutonymphs in a mite species may be of profound evolutionary significance. The phoretic deutonymphs, which attach to the body of the host bee and then drop off in a new nest of that same bee species, insure only the continuation of the same host relationship. But the occurrence of immobile deutonymphs, which remain in the old nest, gives the mite species an opportunity to colonize other species of bees that also nest in abandoned borings.
Parasitism by Ch. krombeini can attain high levels, especially in humid areas of the United States (Bosch & Kemp, 2001). edba5603d18f9fcfb0adbab4449141ba taxon_chaetodactylus_krombeini http://purl.org/dc/dcmitype/Text text/html Management of the Bee Mite Species Chaetodactylus krombeini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Management Inspection of Osmia lignaria nests in semi-translucent paper straws and removal of infested cells is a time-consuming, but effective method to prevent damaging Ch. krombeini infestations in managed colonies. Stripping cocoons out of the nesting materials and managing loose cocoons, instead of whole nests, reduces initial infestation because emerging bees are not forced to walk through infested cells. However, releasing O. lignaria populations as loose cocoons increases dispersal of prenesting females (Bosch and Kemp 2001).
Studies on the Japanese species Chaetodactylus nipponicus, associated with the hornfaced bee, Osmia cornifrons, showed significant reductions of mite infestation in nesting materials treated with endosulfan (60-600 ppm). These same studies indicated that exposure of hornfaced bee nests to high temperatures (e.g., 60 days at 30°C, or three days at 40°C) effectively kills Ch. nipponicus mites without harming hornfaced bees, as long as the bees are in the prepupal stage (Sekita and Yamada 1993, Yamada et al. 1971). Similarly, treating cocoons of O. cornuta and O. rufa during the winter period with a 0.007% solution of endosulfan for a period of 3 min is a very effective method of controlling Ch. osmiae in Europe. It was found that such treatment of cocoons had no negative effect on the bees inside (Krunić et al. 2005). The potential utility of these or similar methods to control Ch. krombeini in Osmia lignaria populations is being tested. 452bfd8f0a421b955dec90e0350002ee taxon_chaetodactylus_lassulus http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus lassulus, phoretic deutonymph, holotype, ex Trichothurgus dubius, Chile, BMOC 96-0916-215 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_lassulus_holotype_BMOC_96-0916-215.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e d58681705ee017bb6fb9090a29e0bbbc taxon_chaetodactylus_lassulus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus lassulus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description (unique character states underlined). Phoretic deutonymph. Longitudinal sclerites on rostral projection (distal extensions of coxal apodemes I) well sclerotized, distinct; rostral parts separated by distinct space and diverging posteriorly. Alveoli ve elongated, situated, outside prodorsal shield. Prodorsal shield small, not extending anteriad of base of se; transversely striated; lateral angles attenuated. Setae si situated outside prodorsal shield, length highly variable from distinctly longer to slightly shorter than 1/2 of distance between their bases. Hysterosomal shield longitudinally striated, except for anterio-lateral part, most lines longer than half of length of hysterosomal shield. No reticulate pattern on hysterosomal shield. Lateral hysterosomal sclerites ventro-lateral, with anterior end situated at anterior level of coxal apodemes III and posterior end at level of posterior apodemes IV. Dorsal setae smooth. Setae c2 situated outside prodorsal shield. Setae c1 and d2 situated outside hysterosomal shield. Setae cp and c2 almost on same transverse level (distance between them usually not exceeding twice the diameter of bases of c2. Setae h3 shorter than femur. Setae h1 and e1 nearly of same length. Coxal setae 1a filiform. Coxal setae 1a, 3a, and 4b situated on soft cuticle. Alveoli of 3a and 4b without surrounding sclerites. Ventral setae 4b distinctly shorter than 3a. Coxal setae 4a almost as long as 3a. Sternal apodeme not bifurcated posteriorly. Posterior apodeme II present, slightly exceeding 1/2 of lateral edge of sternal shield. Proximal and distal extensions of apodemes IV disjunct dorsally. Longitudinal striation between coxae III-IV present. Attachment organ width (including transparent margin) distinctly shorter than distance between 4a. Lateral sclerites of attachment organ not reaching level of 4a. Suckers ad3 (excluding transparent margin) smaller or nearly equal to inner unsclerotized area of suckers ad1+2. Anterior edge of ps2 bases posterior to posterior edge of inner unsclerotized area of suckers ad1+2. Semicircular sclerite distal to base of wa I-II present. Ventral side of claws I-III finely striated. Solenidion σ I shorter than 0.3 of genu I. Setae mG and cG I shorter than combined length of genu and tibia I, both pectinate. Genual setae mG II, pectinate, shorter than combined length of tibia and genu II, longer than mG I, nearly equal to vF II. Femoral setae vF II distinctly shorter than combined length of femur, genu and tibia II. Solenidia ω1 and ω3 distinctly separated, distance exceeds 3 diameters of alveolus of famulus. Tarsal setae la I-II filiform. Tarsal setae wa I-II filiform, slightly widened at base. Genual setae nG III extending end of genu, smooth. Solenidion σ III absent, represented by alveolus. Seta s III subapical. Solenidion φ IV shorter than genu IV. Tarsal setae s and w IV present, uniform in length and width, both shorter than maximum width of tarsus IV. Tarsal setae f and e IV vestigial or absent. Tarsus IV about 3 times longer than its basal width. Other instars unknown. 7c66994e1a95b93d095b1fe9b4131749 taxon_chaetodactylus_lassulus http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Species Chaetodactylus lassulus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription The new species differs from all known species of Chaetodactylus by the autapomorphic characters indicated in the description above (underlined). The relative lengths of setae mG II and vF II (shorter than combined length of femur, genu and tibia II) indicates that the new species belongs to the species group primarily associated with Lithurgus. Beside the autapomorphies, it differs, from all species of this group by the well developed posterior apodemes II, which exceed half of the lateral edge of the sternal shield. The position of setae c1 and d2 outside the hysterosomal shield and the elongation of tarsi I-II (as a result, solenidia ω1 and ω3 become distinctly separated, the distance between them exceeding 3 diameters of alveolus of famulus) are shared with two undescribed species associated with Lithurgus from the New World. Setae d2 (but not c1) are also situated outside the hysterosomal shield in three species associated with Lithurgus (C. ludwigi (Trouessart, 1904), C. dalyi Fain, 1974, and related undescribed species from the Neotropics) and one undescribed species associated with Osmia. Ventral striation of claws I-III is shared with an undescribed species associated with Anthidium from Chile. 668834a4154c3c5165dfdc318046a7bd taxon_chaetodactylus_lassulus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus lassulus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trichothurgus dubius (Sichel, 1867) (type host); Trichothurgus herbsti (Friese, 1905); T. sp. (identified as a new species by Moldenke) 248d46a1c735d67160bf7efeed0b07e8 taxon_chaetodactylus_lithurgi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus lithurgi, phoretic deutonymph, ex Lithurgus apicalis, Idaho, BMOC 96-0510-009 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_lithurgi_paratype_BMOC_96-0510-009.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7a9a0bd5ebf680ee195732c781092cf1 taxon_chaetodactylus_lithurgi http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Chaetodactylus lithurgi on the bee Lithurgus apicalis, Arizona, BMOC 96-0916-191 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_lithurgi_on_host_BMOC-96-0916-191.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b1342c9fd165a4ece04cac37dfe83f70 taxon_chaetodactylus_lithurgi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus lithurgi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, Colorado, Idaho, New Mexico (type locality), Texas. 5cae6d31758a2cdea9db15f0f786c8e2 taxon_chaetodactylus_lithurgi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus lithurgi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgopsis) apicalis Cresson, 1875 (type host); Lithurgus (Lithurgopsis) littoralis Cockerell, 1917; Lithurgus (Lithurgopsis) gibbosus Smith, 1853 57e2d2d886bfa1c833134c35a12c2a2c taxon_chaetodactylus_lithurgi http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus lithurgi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology In Arizona is found phoretic on bee species of the genus Lithurgus flying in the spring. This contrasts to the nearly sympatric species Chaetodactylus abditus associated with Lithurgus species which are active during the fall. 349dc8fd50f226d999a6896f3151dc6a taxon_chaetodactylus_ludwigi http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus ludwigi, phoretic deutonymph, ex Lithurgus scabrosus, New Caledonia, BMOC 91-1015-010 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_ludwigi_BMOC_91-1015-010.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2eb06245f2bd8e48e1bbcb9754860a93 taxon_chaetodactylus_ludwigi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus ludwigi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Federated States of Micronesia: Pohnpei Is. ("Carolines Islands: Ponapé") (type locality);
New Caledonia; French Polynesia: Moorea Is; Indonesia ("Java: Soekaboemi"); India; Madagascar (Fain & Pauly, 2001). c52dc7d7bc7b3bae57969fcff2c86108 taxon_chaetodactylus_ludwigi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus ludwigi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lithurgus (Lithurgus) atratus Smith, 1853 (as Megachile lonalap) (Micronesia) [nest in trunk of Hibiscus ("Hybiscus") with pollen of this plant] (type host) (Trouessart, 1904a) (India, as Lithurgus dentipes, part.)) Fain & Pauly, 2001; Lithurgus pullatus Vachal, 1903 (Madagascar); Lithurgus scabrosus (Smith, 1859) (Java; Moorea Is, near Tahiti; New Caledonia) 6fbc6b075bcef7ba8881542f1545e0d6 taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus melitomae, phoretic deutonymph, ex Melitoma sp., Mexico, BMOC 95-0422-013 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_melitomae_holotype_BMOC_95-0422-013.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2fe4682c87d44041b135e36ad00e0967 taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus melitomae, heteromorphic deutonymph, dorsal view http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_melitomae.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a33310621fb3ea81baaeb3c1cf9fe07b taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/StillImage image/jpeg Heteromorphic deutonymphs of Chaetodactylus melitomae phoretic on Melitoma sp. from Mexico (Campeche) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_melitomae_BMOC-04-0508-299b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e cea79a4277b1bb201860e8139999300a taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/StillImage image/jpeg Heteromorphic deutonymphs of Chaetodactylus melitomae phoretic on Melitoma sp., Honduras BMOC 04-1222-143 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_melitomae_on_host_BMOC-04-1222-143.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 546520964db29a64016b0154c2b4bff6 taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/StillImage image/jpeg Heteromorphic deutonymphs of Chaetodactylus melitomae phoretic on Melitoma sp., Honduras BMOC 04-1222-143 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_melitomae_on_host_BMOC-04-1222-143b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a7119194f8d5332147fe5fc6b2d429b6 taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus melitomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution (show map). Mexico: Campeche, Chiapas, Oaxaca, Tabasco (type locality); Honduras: Yoro. 993fcf193c48eaacdd9a427cbfc5489b taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus melitomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Melitoma marginella, Melitoma segmentaria, Melitoma sp. (type host) (Apidae: Emphorini) (Linsley et al., 1980; our data). Also known from Diadasia sp. (possibly host misidentification or secondary contamination; voucher specimens of the host were not available for study). f140a29c40623375dcd29f47c0c1dbb9 taxon_chaetodactylus_melitomae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus melitomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Linsley et al. (1980) reported that 2.4% and 3.2% of the cells of Melitoma marginella examined at two sites in Chiapas, Mexico, contained heteromorphic deutonymphs of Chaetodactylus sp. nr. ludwigi. Given our collections of C. melitomae from Melitoma spp. from this region, we think Linsley et al. most likely observed this species. Those authors reported that most of these cells contained hundreds of mites, which appeared to have consumed all or substantial amounts of the pollen. The infested cells did not have fragments of bee eggs or larvae suggesting that the egg or young larva is also destroyed by the mites, or that the latter develop successfully only in cells in which oviposition did not take place. Since the deutonymphal stage is dependent upon phoretic transport from the cell in which it develops to a new cell where further development and reproduction occur, burrows of bees which arrange their cells in a series would seem to be suited to these mites. Bees emerging in cells lower down in the series would readily become contaminated if they pass through mite-infested cells (after Linsley et al., 1980). 6299a83b61a8e8d7ea4ba0887c70c187 taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus micheneri, phoretic deutonymph, ex Osmia subaustralis, California, BMOC 03-0514-012 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_micheneri_BMOC_03-0514-012.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f0398e0b8428f62bcfa3dd45d8cdd1a2 taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus micheneri, phoretic deutonymph, holotype http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_micheneri_HDN.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 99d8021945b45a8aafeafbacc8b0c566 taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Chaetodactylus micheneri on a museum specimen of Osmia californica, Utah http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus-micheneri_BMOC-96-0510-111-Osmia_calif.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 0937e401ee623a1c9815e619f92c99dd taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus micheneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description See key. Click here for identification of 3 partially overlapping morphs (based on phoretic deutonymphs only). Known from all stages. Female (unique characters only): Setae c3 not extending beyond posterior level of trochanters IV. Inseminatory canal trumpet-shaped, more than 5 times longer than its width at spermatheca (Fig.). Sclerotized lining layer of outer end of inseminatory canal shorter than 0.4 length of inseminatory canal (Fig.). Setae gT I-II smooth, filiform; hT I smooth, hT II sparsely barbed to almost smooth. Homeomorphic male (unique characters only): Setae d1 not extending beyond e1. Aedeagus folds posterior to dorsal supporting sclerite (Fig.). Transverse processes of dorsal supporting sclerite spirally twisted, band-like (Fig.). Genital valves with posterior cuticular flaps (Fig.). 5b8801bc9aae917c3110264121709dc7 taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus micheneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution
Form 1: USA: California (type locality), Arizona, Colorado, Montana, Utah, Washington, Wyoming; Canada: Alberta.
Form 2: USA: California, Colorado, Idaho, Nevada, Utah, Washington.
Form 3: USA: Michigan; Canada: Northwest Territories.
Unclassified: USA: Oregon. 587f1a3eb92f8f52152d072b952db10f taxon_chaetodactylus_micheneri http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus micheneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Form 1: Osmia (Cephalosmia) subaustralis (type host) (Megachilidae); Form 2: Hoplitis fulgida; O. (Cephalosmia) californica; O. (C.) grinnelli; O. (C.) marginipennis; O. (C.) montana; O. (C. ) subaustralis; Osmia (Acanthosmioides) calcarata; Osmia (Melanosmia) juxta.; Form 3: O. subaustralis.; Unclassified: Osmia montana quadriceps 4291d53cd82bf283e5c572bd40ca1932 taxon_chaetodactylus_nipponicus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus nipponicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan: Honshu (Aomori Pref., Hiraka-machi, Minami-Tsugaru-gun - type locality), Tsushima Is. 5ac0f460f95cd30db65681a72e6c7d03 taxon_chaetodactylus_nipponicus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus nipponicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Osmia) excavata Alfken, 1903 (type host); Osmia (Osmia) cornifrons (Radoszkowski, 1887) ; Osmia (Osmia) taurus Smith, 1873 ; Osmia (Osmia) pedicornis Cockerell, 1919 239602b9b48f746f385a0c278d0d6d00 taxon_chaetodactylus_nipponicus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus nipponicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Pest of Osmia cornifrons managed to pollinate orchards in Japan. 1113f2e2669bdac2cdfd38e0e71b8bba taxon_chaetodactylus_osmiae_ http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus osmiae, phoretic deutonymph, ex Osmia rufa, Germany, BMOC 95-0422-121 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_osmiae_BMOC_95-0422-121.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 834099e2a846c442bee354684c9f2a82 taxon_chaetodactylus_osmiae_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus osmiae ( http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution France (type locality); Belgium, England, Spain, Germany, Hungary, Croatia, Romania (Klimov & OConnor, 2008) (because historical literature may not distinguish this species from Ch. zachvatkini, only verified records are listed here, see synonymy for additional localities) 9cd23949f534970f904f9a90098d323a taxon_chaetodactylus_osmiae_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus osmiae ( http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia rufa (as Osmia bicornis and Osmia fronticornis) (France) (type host); Osmia cornuta (Latreille, 1805); Osmia niveata (Fabricius, 1804); Osmia tricornis Latreille, 1811 ; ; (because historical literature may not distinguish this species from Ch. zachvatkini, only verified records are listed here, see synonymy for additional hosts) 8500ed325738584c519869a610fa7116 taxon_chaetodactylus_osmiae_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus osmiae ( http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Serious pest of managed populations of Osmia cornuta and O. rufa in Europe (Bosch, 1992; Krunić et al., 2005). 969852cc4301bd50cef4ca9698a2a79b taxon_chaetodactylus_reaumuri http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus reaumuri, phoretic deutonymph, ex Osmia brevicornis, Poland, BMOC 04-0508-019 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_reaumuri_BMOC_04-0508-019.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 372c6e94826c6906e9b00fd404f03253 taxon_chaetodactylus_reaumuri http://purl.org/dc/dcmitype/StillImage image/jpeg Heteromorphic deutonymphs of Chaetodactylus reaumuri phoretic on Osmia brevicornis, Poland http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_reaumuri_on_host_BMOC_04-0508-001c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8270393f31ed08493bbbcaa7928f1eb1 taxon_chaetodactylus_reaumuri http://purl.org/dc/dcmitype/StillImage image/jpeg Heteromorphic deutonymphs of Chaetodactylus reaumuri phoretic on Osmia brevicornis, Poland http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_reaumuri_on_host_BMOC_04-0508-001a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f751a644bae6a299a40c68bc030b5e04 taxon_chaetodactylus_reaumuri http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus reaumuri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Czech Republic: Kolín ("Kolin, Bohemen") (type locality), Odry "Odran, Oostenrijksch Silezië" (Oudemans, 1905b); Germany; Ukraine: Crimean Peninsula; Georgia (Zachvatkin, 1941); Greece: Corfu (Türk & Türk, 1957). d5da4925aa426ca4d0780a5ee7d99c24 taxon_chaetodactylus_reaumuri http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus reaumuri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia niveata (Fabricius, 1804) (as "Osmia rufiventris Panz." (type host) probably misspelled for Osmia fulviventris (Panzer, 1798); Osmia brevicornis (Fabricius, 1798) (as Osmia panzeri); Osmia brevicornis leucogastra Morawitz, 1875 (as Osmia leucogastra); Stelis phaeoptera murina Pérez, 1884 (as Stelis murina) (cleptoparasitic on Osmia, including O. niveata) ; (Oudemans, 1905b; Zachvatkin, 1941) 30c4954b0a9f3b3f21f505b414d2e929 taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus rozeni, phoretic deutonymph, ex Osmia georgica, North Carolina, BMOC 04-0508-007 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_rozeni_holotype_BMOC_04-0508-007.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 795beede64b3a64b5806a685feea558c taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus rozeni http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_rozeni.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e be37fa367966cb54b2ae5a0da86050ff taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Chaetodactylus rozeni on a museum specimen of Osmia georgica, Texas http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus-rozeni_BMOC-04-0508-009c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 63067cce1be0f113e854dac376a7172f taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus rozeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Krombein (1962) reported heteromorphic deutonymphs of Chaetodactylus associated with Osmia caerulescens from New York. The deutonymphs were similar to those of Ch. krombeini except for having slightly but consistently shorter body setae (Krombein, 1962). Chaetodactylus rozeni has dorsal setae that are distinctly shorter than in Ch. krombeini, however, these two species also differ in many other morphological details. Because O. caerulescens and O. georgica belong to the same subgenus, Helicosmia, and their ranges are overlapping, we believe that Krombein (1962, 1967) probably dealt with mites identical with Ch. rozeni. b0c3a3dd136dc5bafd36636e929973d6 taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus rozeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: North Carolina (type locality), Tennessee (see also notes), Texas. fb35ac7346675ac935e903314f733495 taxon_chaetodactylus_rozeni http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus rozeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia (Helicosmia) georgica (Megachilidae) (type host) (see also notes). Collected from Chrysis coerulans (Hymenoptera: Chrysididae), a polyxenous cleptoparasite of wasps and bees. 0bd182bc56fb2d02bf5f861da3e7b416 taxon_chaetodactylus_zachvatkini http://purl.org/dc/dcmitype/StillImage image/jpeg Chaetodactylus zachvatkini, phoretic deutonymph, ex Osmia tricornis, Italy, BMOC 03-0523-005 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Chaetodactylus_zachvatkini_holotype_BMOC_03-0523-005.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7561c4d567e788bcdbde5bf786a789de taxon_chaetodactylus_zachvatkini http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Chaetodactylus zachvatkini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. This species was described and depicted by Zachvatkin (1941) as Chaetodactylus osmiae (Dufour). Van Asselt (2000) redescribed Chaetodactylus osmiae from Belgium (which is very close to the type locality) and from the type host, Osmia rufa. He believed that Zachvatkin had the same species but erroneously depicted its dorsum without prodorsal shield (actually it is present on the figure and mentioned in the description and in the key) and short dorsal setae (longer in true Ch. osmiae). Because Zachvatkin describes Ch. osmiae as having "lateral suckers [=ps2] situated on the same transverse level with central ones [ad1+ad2]" (as in true Ch. osmiae) but figures mites with shorter setae, we believe that he dealt with two very similar species and one of them, having "shorter" setae, is new, Chaetodactylus zachvatkini Klimov and OConnor, 2008. Unfortunately, we could not find any specimens that match Ch. osmiae sensu Zachvatkin, 1941 or even true Ch. osmiae in his collection in Zoological Institute of the Russian Academy of Sciences, Saint Petersburg (ZIN). Material from Italy (including the holotype) was collected by Dr. S. V. Mironov in the insect collection of ZIN on our request. Since insect-associated mites described by Zachvatkin (1941) were collected in ZIN and this author mentioned that Ch. osmiae occurs in Italy on Osmia tricornis (p. 398), we believe that our mites originated from the same bee hosts that were sampled by Zachvatkin. f90e0be65c618a0e8a22839f64052cc7 taxon_chaetodactylus_zachvatkini http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Chaetodactylus zachvatkini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Italy: Liguria, San Remo (type locality); Spain; Tunisian Republic. 04e641ae60b0ce87d41528f46c6f4a1f taxon_chaetodactylus_zachvatkini http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Chaetodactylus zachvatkini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Osmia tricornis Latreille, 1811 (type host); Osmia cornuta (Latreille, 1805); Osmia latreillei (Spinola, 1806); Osmia niveata (Fabricius, 1804) 9d515c681368f955567f061862f614d3 taxon_chaetodactylus_zachvatkini http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Chaetodactylus zachvatkini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology May be a pest of managed Osmia colonies in southern Europe and northern Africa. a7ebca8e56964a4aa126e4cb217775c1 taxon_cheletophyes http://purl.org/dc/dcmitype/StillImage image/jpeg Cheletophyes panamensis, female, ex nest of Xylocopa frontalis, Costa Rica, BMOC 03-1228-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Cheletophyes_panamensis_f_BMOC_03-1228-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2038c767dba5e3702289360beef6b049 taxon_cheletophyes http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Cheletophyes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus Cheletophyes is distributed in the Oriental, Afrotropical, and Neotropical regions, with 13 species obligatory associated with large carpenter bees, genus Xylocopa (several records from other hosts are probably accidental) (Fain and Bochkov, 2001a,b; Klimov at al., 2006). All stages of these mites are predators of different microarthropods feeding on provision in the bees' nests (Eickwort, 1994). Females of Cheletophyes disperse in mesosomal acarinaria of adult bees. According to the hypothesis of OConnor (1993), the relationships between Cheletophyes and their bee hosts are mutualistic, and the bees have developed mesosomal acarinaria to transfer these predacious mites controlling nest cleptoparisites. Our study (Klimov et al., 2006) suggests that despite the fact that the mites do not directly depend on resources provided by the bees, they are highly specific to their hosts. One species, Cheletophyes panamensis, was recorded for North America (Klompen et al., 1984). This species was found in the nest of its host only. Neotropical large carpenter bees lack an acarinarium. So far, phoretic specimens of Ch. panamensis were not discovered on their host's bodies despite our significant effort to find them. 6e85ff859703b626ef35d8756939d270 taxon_cheletophyes_panamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Cheletophyes panamensis, female, ex Xylocopa frontalis, Costa Rica, BMOC 03-1228-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Cheletophyes_panamensis_f_BMOC_03-1228-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b8f53773946184c7f94b914b16c7614a taxon_cheletophyes_panamensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Cheletophyes panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Panama (type locality), Costa Rica (Klompen et al., 1984; our data). d402af6f18ec6f623f5d9748e25ceaf3 taxon_cheletophyes_panamensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Cheletophyes panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Megaxylocopa) frontalis (Olivier, 1789) (Klompen et al., 1984). e190f22f0358faca80de2c30700635b1 taxon_cheletophyes_panamensis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Cheletophyes panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology The species was originally found in the nest of Xylocopa frontalis constructed in the tree Luehea seemannii Triana & Planch., 1862 (Malvales: Tiliaceae). c8b056d6551c3964d0f21c04d195c02d taxon_cheyletidae http://purl.org/dc/dcmitype/StillImage image/jpeg Cheletophyes panamensis, phoretic female, nest of Xylocopa frontalis, Costa Rica, BMOC 03-1228-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Cheletophyes_panamensis_f_BMOC_03-1228-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 57c8be4858aa566aeef47261a5a09dfb taxon_cheyletidae http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic females on the propodeum of the bee Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_on_bee_e_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 00cdc8504974ff16c645b6c9cc1c527b taxon_cheyletidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Cheyletidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This cosmopolitan family includes 75 genera and 438 described species (Bochkov and Fain, 2001a; Zhang et al., 2011). Some species are predators occurring in soil, forest litter, under tree bark and on foliage, in spore tubes of polypore fungi, nests of birds, mammals or insects, quills of birds, grain storages, and house dust. Several species are ectoparasites of different terrestrial vertebrates. Females of some cheyletid genera were found phoretic on different insects, including bees: Cheletophyes Oudemans, 1914 on Xylocopa spp., Bak furcatus on Apis cerana (Hymenoptera: Apidae), Nodele (Aztecocheyletus) on Aztecanthidium tenochtitlanicum (Hymenoptera: Megachilidae), Samsinakia on Coleoptera: Tenebrionidae, Hypopicheyla Volgin, 1969 on Aradus (Hemiptera: Aradidae) and Tenebrionidae, Cheyletia Haller, 1884, Cheletophanes Oudemans, 1904, and Cunlifiella Volgin, 1964 on Aradus, and Paracaropsis Volgin, 1969 on Laphria and Pagidolaphria (Diptera: Asilidae) (Bochkov and Fain, 2001a; Bochkov and Mironov, 1998; Klimov, 1997; Volgin, 1969; our data).
While associations of Cheletophyes and Nodele (Aztecocheyletus) conquistador with their bee hosts are obligatory, many other cheyletid taxa reported from bees possibly represent opportunistic species, occupying a broad range of habitats including bee nests. The list below gives information on species of Cheyletidae recorded from bees.


Acaropsella sp. (Apis mellifera, Czech Republic)
Acaropsis sollers Kuzin, 1940 (Apis mellifera, Czech Republic, Kyrgyzstan)
Bak furcatus Gerson and Fain, 1991 (Apis cerana, Tailand, Philippines)
Cheletomimus bregetovae (Volgin, 1969) (Apis cerana, Philippines)
Cheletomorpha lepidopterorum (Shaw, 1794) (Apis mellifera, Germany, Russia)
Cheletophyes apicola Fain, Lukoschus and Nadchatram, 1980 (Xylocopa latipes, Xylocopa pentachroma, Malaysia, Celebes, Philippines)
Cheletophyes clavipilis Fain, Lukoschus et Nadchatram, 1980 (Xylocopa latipes, Malaysia)
Cheletophyes decorus Bochkov and Klimov, 2004 (Xylocopa tranquebarica (Fabricius, 1804), Vietnam)
Cheletophyes eckerti Summers and Price, 1970 (probable synonyms: Ch. harnaji Putatunda & Kapil, 1988; Ch. haryanaensis Putatunda & Kapil, 1988; Ch. newtoni Putatunda & Kapil, 1988; Ch. orientalis Putatunda & Kapil, 1988; Ch. ruttneri Putatunda & Kapil, 1988; Ch. shendei Putatunda & Kapil, 1988) (Xylocopa aestuansIndia (Xylocopa imitatortera in Angola, bees in Congo, and Bembix ugandensis in Congo after Fain & Bochkov, 2001))
Cheletophyes indiacus Smiley and Whitaker, 1981 (=Ch. xylocopae Ramaraju et Mohanasundaram, 1999) (Xylocopa sp, India)
Cheletophyes mbomba Bochkov, Klimov and OConnor, 2006 (Xylocopa torrida, Africa)
Cheletophyes occisor Bochkov and Klimov, 2004 (Xylocopa aruana, Papua New Guinea)
Cheletophyes panamensis Klompen, Méndez and Lukoschus, 1984 (Xylocopa frontalis, Costa Rica, Panama)
Cheletophyes sp. (Xylocopa, Australia)
Cheletophyes torridae Bochkov, Klimov and OConnor, 2006 (Xylocopa torrida, Africa)
Cheletophyes venator (Vitzthum, 1920) (=Ch. aurorae Haitlinger 2000) (Xylocopa nigrita, Africa)
Cheletophyes vespae Fain and Bochkov, 2001 (Vespa tropica (accidental host, should be associated with Xylocopa), Malaysia)
Cheletophyes vitzthumi Oudemans, 1914 (Xylocopa caffra, South Africa)
Cheyletus carnifex Zachvatkin, 1935 (Apis mellifera, Czech Republic)
Cheyletus eruditus Schrank, 1781 (Apis mellifera, Bombus, Czech Republic, Poland, former USSR, India, New Zealand)
Cheyletus malaccensis Oudemans, 1903 (Apis, India)
Eucheyletia bakeri Volgin, 1963 (Bombus, USA)
Eucheyletia flabellifera (Michael, 1878) (Apis mellifera, Czech Republic)
Nodele (Aztecocheyletus) conquistador Bochkov and Klimov, 2004 (Aztecanthidium tenochtitlanicum, Mexico)
(Banaszak 1980, Bochkov and Klimov 2004, Chmielewski 1972, Chmielewski 1991, Corpuz-Raros 1988, Corpuz-Raros 2000, Fain and Bochkov 2001a, Gerson and Fain 1991, Grobov 1975, Haragsim et al. 1978, Homann 1933, Klimov et al. 2006, Klompen et al. 1984, Kumar and Kumar 1995, Macfarlane 2005, Malabanan and Corpuz-Raros 1998, Tomaszewska 1988, Walter et al. 2002)
467dd26e28b429e99a0cbb618c4f1bf8 taxon_diadasiopus http://purl.org/dc/dcmitype/StillImage image/jpeg Diadasiopus eickworti, phoretic deutonymph, ex Diadasia opuntiae, USA: California, BMOC 91-0402-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Diadasiopus_eickworti_BMOC_91-0402-002_holotype.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a2844cdb982357751a8d3f3fc16970ae taxon_diadasiopus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Diadasiopus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Known from phoretic deutonymphs, are adults collected but not described. Molecular phylogenetic analysis calls for reconsidering morphology-based placement of this genus in Horstiinae. The following apomorphic character states joining Diadasiopus with Horstiinae should be re-evaluated: the rounded subcapitular remnant and striate pattern on the dorsal idiosoma (OConnor, 1997). 5156e080c0ffc06c67d2032c7289c1d0 taxon_diadasiopus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Diadasiopus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution New World. e354d97cb8b9213d002eecf359f29c84 taxon_diadasiopus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Diadasiopus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Associated with bees of the genus Diadasia and rarely Anthophora (Apidae) (OConnor, 2001). e8c15c65519e0e32f41b36907ebe1b56 taxon_ereynetes http://purl.org/dc/dcmitype/StillImage image/jpeg Ereynetes boharti (Ereynetidae) from nest of Nomia melanderi (modified from Hunter and Cross, 1968) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Ereynetes_boharti.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 96953a274a455f37d0b41887f45f5a2d taxon_ereynetes http://purl.org/dc/dcmitype/StillImage image/jpeg Ereynetes (Anereynetes) meliponae, female, ex nest of Melipona scutellaris (modified from Flechtmann et al., 1985) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Ereynetes_meliponae_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bc8368472093ba34c6077c0478336b1d taxon_ereynetes http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Ereynetes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes five subgenera and 51 species free-living predators that live in mosses, lichens, leaf litter, bat guano, dung, scarabaeid beetles, nests of mammals and birds, rotting wood, galleries of bark beetles and under bark (Fain and Camerik, 1994). Two subgenera and two species have been found in nests of bees. b5287e7cc53b53482350a40790135ca1 taxon_ereynetes_ereynetes_boharti http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Ereynetes (Ereynetes) boharti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Wyoming. 8146f44a918dfd37cf2d07a07f030ec3 taxon_ereynetes_ereynetes_boharti http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Ereynetes (Ereynetes) boharti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Nomia melanderi Cockerell, 1906 (Halictidae). 0edcee8997de37bde09f1750ec3230d3 taxon_ereynetes_ereynetes_boharti http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Ereynetes (Ereynetes) boharti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species has been collected only once from artificial nests of the bee Nomia melanderi (Halictidae) containing many dead prepupae and mites of the genera Sancassania (Acaridae), Anoetus (Histiostomatidae), and the family Rhodacaridae. Probably, the normal habitat of this species is soil surface and the mites are able to invade the artificial nests because of relatively free access to the cells made possible by the type of nest construction (Hunter and Cross, 1968). The species probably is a predator, like others representatives of the genus Ereynetes. c2fbe670fb940a9141a615f0f30be544 taxon_ereynetes_anereynetes_meliponae http://purl.org/dc/dcmitype/StillImage image/jpeg Ereynetes (Anereynetes) meliponae, female, ex nest of Melipona scutellaris (modified from Flechtmann et al., 1985) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Ereynetes_meliponae_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f1a8b3f1586da4a7ef4470390c10869a taxon_ereynetes_anereynetes_meliponae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Ereynetes (Anereynetes) meliponae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Pernambuco. b726c8243d6265199ede4908e538dc2a taxon_ereynetes_anereynetes_meliponae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Ereynetes (Anereynetes) meliponae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Melipona scutellaris Latreille, 1811 (Apidae: Meliponini). d838586c35be340a450df0bdf1332023 taxon_ereynetes_anereynetes_meliponae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Ereynetes (Anereynetes) meliponae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected once in the nest of Melipona scutellaris. Larvae and prelarvae were found inside females. aa69b22b5779813655e55ff4a76a883e taxon_ereynetidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Ereynetidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes 29 genera and 180 species grouped in three subfamilies (Kethley, 1982; Zhang et al., 2011). Species of Erynetinae (Ereynetes) are free-living predators in mosses, lichens, leaf litter, and under bark. Two species of Ereynetes were found in bee nests but it is not clear if these associations are obligate. Other ereynetines, Riccardoella Berlese, 1923, are obligate pulmonary parasites of terrestrial slugs, or in the genus Hydranetes, subelytral parasites of aquatic beetles. All stages, except the prelarva, are mobile in Ereynetinae. The subfamily Lawrencarinae includes parasites of the nasal passages of frogs and toads. The tritonymph was thought to be missing in the Lawrencarinae (Fain, 1972). The missing nymph of Lawrencarinae was, however, discovered later (André and Fain, 2000). The subfamily Speleognathinae contains nasal parasites of birds and mammals. In the speleognathine development, the three nymphal stages are calyptostatic, without any structures other than pharynx and the larva gives rise to the adult (Fain, 1972). efba0f39d1217c27b7cb7fcbf0a64ac6 taxon_erythraeidae http://purl.org/dc/dcmitype/StillImage image/jpeg Leptus ariel (Erythraeidae), larva, dorsal view (modified from Southcott, 1989) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Leptus_ariel.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e fd1ef21bf2f54aad3f098b9275b4265f taxon_erythraeidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Erythraeidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes 55 genera and 771 species (Vercammen-Grandjean, 1973; Zhang et. al., 2011). Larvae of many these genera are ectoparasites of insects, millipedes, or arachnids, whereas postlarval instars are predatory. One genus, Leptus, occurs on bees. In addition, an unidentified genus was recorded from the Philippines from Apis cerana (Malabanan and Corpuz-Raros, 1998). 95feb2d5f0cfba70a42bb92749c22fd3 taxon_eucheyletia http://purl.org/dc/dcmitype/StillImage image/jpeg Eucheyletia bakeri Volgin, 1963, female, dorsal view (left) and gnathosoma, dorsal view (right),   bumblebee nets, Alaska (modified from Baker, 1949) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Eucheyletia_bakeri.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3194d5720c41e6d8ed07de43ce22a12a taxon_eucheyletia http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Eucheyletia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes ten species distributed in the Holarctic and Oriental regions. It is primarily associated with nests of rodents. Species of this genus were also found in nests of other small mammals and birds, on oat seeds, barn and house dust (Fain and Bochkov, 2001a; Volgin, 1969). Two species were found in associations with bees. Eucheyletia bakeri is known from a single record from the nest of a bumblebee in the USA (Baker, 1949; Volgin, 1963). Eucheyletia flabellifera was collected from honeybee hives in Europe (Haragsim et al., 1978). ea957fc2216d2803a52d3fa07d6e4385 taxon_eucheyletia_bakeri http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Eucheyletia bakeri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Alaska, Beaver Mountains (Baker, 1949). f75b383234782d731f4e035a12a66c6f taxon_eucheyletia_bakeri http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Eucheyletia bakeri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Bombus sp. (Baker, 1949). 6fa64fde87cbbcd2a11bd0fbb2ff5269 taxon_eucheyletia_bakeri http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Eucheyletia bakeri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Three females of this species have been collected from the nest of a bumble bee (Baker, 1949). This record is probably accidental because it has not been confirmed and this type of habitat is unusual for the genus. On the other hand, there are mite genera (Kuzinia, Parasitellus) that occur both in rodent and bumble bee nests. This may be explained by the fact that bumble bees often construct their nests in borrows of rodents.
85f86663cd6986bf8397a0d578d4365d taxon_eumellitiphis http://purl.org/dc/dcmitype/StillImage image/jpeg Eumellitiphis mellitus, male, ex Lestrimelitta limao, Panama, BMOC 88-0729-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Eumellitiphis_mellitus_BMOC_88-0729-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a4794b7e7c4f27a01c44539b24b03a06 taxon_eumellitiphis http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Eumellitiphis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes three named species inhabiting nests of stingless bees (Apidae: Meliponini) in the Neotropical and Oriental regions (Delfinado-Baker and Baker, 1988; Turk, 1948). Eumellitiphis mellitus Turk was found in the nests of Lestrimelitta limao in Trinidad and Tobago, Panama, and Brazil. Eumellitiphis inouei Delfinado-Baker et Baker from Geniotrigona thoracica and Lophotrigona canifrons from Indonesia. Eumellitiphis philippinensis Delfinado-Baker et Baker was found in nests of an unidentified species of Trigona from the Philippines (Turk, 1948; Delfinado-Baker and Baker, 1988; our data). b8b542ce22c1c8f6d0e4e190c34ac302 taxon_eumellitiphis_inouei http://purl.org/dc/dcmitype/StillImage image/jpeg Eumellitiphis inouei, female, ex Geniotrigona thoracica, Indonesia (Sumatra) (modified from Delfinado-Baker and Baker, 1988). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Eumellitiphis_inouei.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 038e410bdda2aaf0749f71504bcf6012 taxon_eumellitiphis_inouei http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Eumellitiphis inouei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Sumatra (type locality: Sumatera Barat, Sawahlunto); Malaysia. eb61db2678c80caf0aa5df886ccf2a5d taxon_eumellitiphis_inouei http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Eumellitiphis inouei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Geniotrigona thoracica (Smith, 1857) (as Trigona thoracica) (type host); Lophotrigona canifrons (Smith, 1857) (as Trigona canifrons); Trigona sp. c5eacfbb420b6eb5561c857bccf34f04 taxon_eumellitiphis_mellitus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Eumellitiphis mellitus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. This species has been originally described (protonymph, male, female) from nests of Lestrimelitta limao from Trinidad (Turk, 1948). Delfinado-Baker and Baker (1988) found a female and protonymph in the nest of the same host and included a figure of the female. The authors also mentioned that they have compared their material with the Turk's types. We have a male that has the dorsal shield very similar to that of the female figured by Delfinado-Baker and Baker (1988). However, the anterior part of spermatodactyl in our male is slightly S-shaped, but it is straight in the Turk's figure (Fig. 45). Thus, our specimens probably are not authentic Eumellitiphis mellitus. cfed184953db3e52a362e4e786e0695f taxon_eumellitiphis_mellitus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Eumellitiphis mellitus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Trinidad and Tobago: Arima (type locality), Brazil (no additional data); Panama: Colón (Turk, 1948; Delfinado-Baker and Baker, 1988; our data). 32584802a8818d62d01bf107558482ef taxon_eumellitiphis_mellitus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Eumellitiphis mellitus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lestrimelitta limao (Apidae) (type host). This mite species might actually have a wide range of hosts, because Lestrimelitta robs provision in the nests of other stingless bees (Michener, 2000). 05844dfbdec03aee486237b42fc43a1f taxon_eumellitiphis_philippinensis http://purl.org/dc/dcmitype/StillImage image/jpeg Eumellitiphis philippinensis, female, ex "Trigona" sp. nest, Philippines (modified from Delfinado-Baker and Baker, 1988) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Eumellitiphis_philippinensis.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7683d123a7cb47925ed33b856bfc0baa taxon_eumellitiphis_philippinensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Eumellitiphis philippinensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Philippines: Leyte (type locality: Inopacan). 679bc6117ff61808cfa4bbe8480faef5 taxon_eumellitiphis_philippinensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Eumellitiphis philippinensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona sp. (nest) 4efa476c2fdb9e59db097d918e1c7f59 taxon_horstiella http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella quadrata, holotype, phoretic deutonymph, ex Epicharis rustica, Guyana, BMOC 82-0124-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_quadrata_holotype_BMOC_82-0124-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b7b54225f9dfc01d4986e6e8b122d31b taxon_horstiella http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Genus Horstiella http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription Phoretic deutonymph (after Ochoa and OConnor, 2000). Dorsum with faint to well-developed pattern of lines, transverse on propodosoma and anterior hysterosoma, longitudinal on posterior hysterosoma. External vertical setae ve absent, represented by pair of vestigial alveoli posterolateral to internal verticals (vi). Supracoxal setae scx absent. Venter with coxal apodemes well developed, coxal fields III-IV closed; coxal fields III separated, contiguous or rarely fused medially, coxal fields IV closed. Coxal setae 1a, 3a, 4b absent, coxal setae 4a in the form of large, striate conoids. Genital papillae asymmetrical, with posterior pair more attenuate. Attachment organ with anterior and median suckers normally formed, with two pairs of striate conoids and five large cuticular suckers. Legs normally formed, tarsal setation 8-8-8-8; setae aa I, ba I-II, and nG III absent; solenidion ω2 absent from tarsus I, represented by vestigial alveolus, solenidion σ absent from genu III. All pretarsi with hooked empodial claws arising directly from tarsal apices.
Feeding stages unknown. 91f849c53d5d6e2d2f2c922f15e37850 taxon_horstiella http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Horstiella http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Neotropical region, ranging from central Mexico through Brazil (Ochoa and OConnor, 2000). f46096a95495719e58dc2350cf21ccb0 taxon_horstiella http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Horstiella http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Species of Horstiella are exclusively associated with the ground nesting bee genus Epicharis in the Neotropics. The mites occur on the subgenera Triepicharis, Pareipcharis, Hoplepicharis, Epicharis (s.str.), and Epicharana, while the subgenera Epicharoides and Epicharitides lack these mites. The association of Horstiella and Epicharis is quite unusual in that most species of Epicharis harbor two species of Horstiella, a condition known as synhospitaly (Ochoa and OConnor, 2000). The mite species pairs were almost always spatially segregated on an individual host. No host specificity was detected, suggesting that this association is shaped after the initial diversification of the genus Epicharis (Ochoa and OConnor, 2000).
Most of Horstiella species are commonly found on the bee's propodeum, while Horstiella megamyzidos specifically attaches under the lateral edges of the metasomal tergites and under the sternites. In male bees, the more frequent attachment of mites to the ventral region of the mesosoma and metasoma correlates with the mating position of the host bees. Because only the female bee makes nests, mites developing in a cell with a male bee would normally have no opportunity to found new colonies. Migrating to the ventral surface of male bees would, therefore, give the mites a selective advantage because this can facilitate transfer onto the body of a female bee during copulation, which occurs with the male above the female (Ochoa and OConnor, 2000).
Phylogeny of Horstiella, Medeus, and Horstia based on morphology was presented by Ochoa and OConnor (2000). 8a2dca7e8eebb30b581a2f557f13e00f taxon_horstiella_armata http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella armata, phoretic deutonymph, ex Epicharis rustica, Guyana, BMOC 82-0124-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_armata_BMOC_82-0124-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 283466476ccc7f5c189c904d87441b98 taxon_horstiella_armata http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella armata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Trinidad and Tobago: Trinidad, St. Augustine (type locality); Costa Rica; Panama, Guyana; French Guiana, Suriname, Colombia, Ecuador, Brazil, Mexico, Nicaragua (Ochoa & OConnor, 2000) 5c254ed035bd24f619bc3bffca81103f taxon_horstiella_armata http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella armata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Eulaema (Apeulaema) cingulata (Fabricius, 1804) (as "Euglossa fasciata Lep.") (type host, misidentification, see Ochoa & OConnor, 2000); Epicharis (Epicharana) angulosa Snelling, 1984 (Costa Rica); Epicharis (Epicharana) bova Snelling, 1984 (Costa Rica); Epicharis (Epicharana) elegans Smith, 1861 (Mexico, Nicaragua); Epicharis (Epicharana) flava (Friese, 1900) (Ecuador); Epicharis (Epicharana) rustica (Olivier, 1789) (Costa Rica, Panama, Guyana, Suriname, Colombia, Ecuador, Brazil); Epicharis (Epicharis) umbraculata (Fabricius, 1804) (French Guiana, Brazil); after Ochoa & OConnor, 2000 61a8b0839dafbb89f861173108574d06 taxon_horstiella_armata http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella armata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella quadrata on the same host, but topologically segregated from it. 1478912c6d4c2648b199e4a679d024d0 taxon_horstiella_concentrica http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella concentrica, phoretic deutonymph, holotype, ex Epicharis metatarsalis, Costa Rica, BMOC 97-0331-015 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_concentrica_holotype_BMOC_97-0331-015.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e aa4cc9f28d3e60ef3b2639c13f07fe1f taxon_horstiella_concentrica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella concentrica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Costa Rica: Heredia (Finca La Selva, near Puerto Viejo) (type locality); Guyana; Bolivia (Ochoa & OConnor, 2000) 890f832c2cd311d82f1b45f3e7665fc9 taxon_horstiella_concentrica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella concentrica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Parepicharis) metatarsalis Friese, 1899 (type host) (Costa Rica); Epicharis (Parepicharis) zonata Smith, 1854 (Guyana, Bolivia); after Ochoa & OConnor, 2000 0e9ca2c9d6ab0e708248b46472d69ce8 taxon_horstiella_concentrica http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella concentrica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella variabilis on the same host, but topologically segregated from it. 9f17a376706869ed14479d426fda1199 taxon_horstiella_megamyzidos http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella megamyzidos, phoretic deutonymph, holotype, ex Epicharis fasciata, Brazil, BMOC 96-0916-039 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_megamyzidos_holotype_BMOC_96-0916-039.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e cd6f837b7efbf3ff5bc685b1bb49b3d1 taxon_horstiella_megamyzidos http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella megamyzidos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Pará: Belém (type locality); Venezuela; Costa Rica; Panama; Guyana; Colombia; Peru (Ochoa & OConnor, 2000) ec97896e3a147b8b21a7e9f9eb6bc141 taxon_horstiella_megamyzidos http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella megamyzidos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Hoplepicharis) fasciata Lepeletier and Audinet-Serville, 1828 (type host) (Brazil, Venezuela, Cost Rica); Epicharis (Hoplepicharis) monozona Mocsáry, 1899 (Panama); Epicharis (Hoplepicharis) affinis Smith, 1874 (Guyana, Colombia, Peru, Brazil); after Ochoa & OConnor, 2000 a0138b904ef8e8dfad52090d786bfdc7 taxon_horstiella_megamyzidos http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella megamyzidos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella snellingi on the same host. Commonly found under the edges of metasomal tergites. 8a609f9199c6c7b601c5960d4030d9bc taxon_horstiella_mourei http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella mourei, phoretic deutonymph, holotype, ex Epicharis analis, Brazil, BMOC 96-0916-042 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_mourei_holotype_BMOC_96-0916-042 http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 82cacea142561f6bc64a56b75b3da1d1 taxon_horstiella_mourei http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella mourei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Goias: Aragarcas (type locality) (Ochoa & OConnor, 2000) 01c0202ea66c7e8d2b091a7a6fb1b8b2 taxon_horstiella_mourei http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella mourei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Triepicharis) analis Lepeletier, 1841 (part. as E. schrottkyi) (type host) (Brazil) (Ochoa & OConnor, 2000) c7f80605c6dbcb52d9f213fae8ba8c8c taxon_horstiella_quadrata http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella quadrata, phoretic deutonymph, holotype, ex Epicharis rustica, Guyana, BMOC 82-0124-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_quadrata_holotype_BMOC_82-0124-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c185f3ef4b0690dd62e16cd6ec9d3908 taxon_horstiella_quadrata http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella quadrata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Guyana: Cuyuni-Mazaruni (Bartica District, Penal Settlement) (type locality); Costa Rica; Panama; Trinidad and Tobago; Suriname; Colombia; Ecuador; Peru; Brazil; Mexico; Nicaragua. 96296d4f810446d65a571181910c6afb taxon_horstiella_quadrata http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella quadrata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Epicharana) rustica (Olivier, 1789) (type host) (Guyana, Costa Rica, Panama, Trinidad and Tobago, Suriname, Colombia, Ecuador, Peru, Brazil); Epicharis (Epicharana) bova Snelling, 1984 (Costa Rica); Epicharis (Epicharana) elegans Smith, 1861 (Mexico, Nicaragua, Costa Rica); Epicharis (Epicharis) bicolor Smith, 1854 (Brazil); Epicharis (Epicharis) umbraculata (Fabricius, 1804) (Brazil); ; after Ochoa & OConnor, 2000 c1af161d70c67c10e37f1bd8fecf00fa taxon_horstiella_quadrata http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella quadrata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella armata on the same host, but topologically segregated from it. 1f7fc6d42c143bc423aa655a80e15ba3 taxon_horstiella_snellingi http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella snellingi, phoretic deutonymph, holotype, ex Epicharis affinis, Guyana, BMOC 95-0310-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_snellingi_holotype_BMOC_95-0310-002 http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b03125ca5420debdb605728d65b5b57e taxon_horstiella_snellingi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella snellingi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Guyana: Cuyuni-Mazaruni (Bartica District, Penal Settlement) (type locality); French Guiana; Trinidad and Tobago(Trinidad); Venezuela; Colombia; Peru; Brazil; Costa Rica; Panama (Ochoa & OConnor, 2000) a1c99803c383f98c58e3d33c8deed3e6 taxon_horstiella_snellingi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella snellingi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Hoplepicharis) affinis Smith, 1874 (type host) (Guyana, French Guiana, Colombia, Peru, Brazil); Epicharis (Hoplepicharis) fasciata Lepeletier and Audinet-Serville, 1828 (Trinidad and Tobago, Venezuela, Brazil); Epicharis (Hoplepicharis) lunulata Mocsáry, 1899 (Costa Rica); Epicharis (Hoplepicharis) monozona Mocsáry, 1899 (Panama); after Ochoa & OConnor, 2000 722d8e1b3ceb3935591682db2de69cb6 taxon_horstiella_snellingi http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella snellingi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella megamyzidos on the same host. Commonly found on the propodeum, wing bases, legs. 0646572e9e9186df343afa70bfe600a3 taxon_horstiella_variabilis http://purl.org/dc/dcmitype/StillImage image/jpeg Horstiella variabilis, phoretic deutonymph, holotype, ex Epicharis metatarsalis, Costa Rica, BMOC 97-0331-015 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Horstiella_variabilis_holotype_BMOC_97-0331-015 http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ff63715b570ed1063fd8f4df225d35b3 taxon_horstiella_variabilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Horstiella variabilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Costa Rica: Heredia (Finca La Selva: near Puerto Viejo) (type locality); Guyana; Bolivia (Ochoa & OConnor, 2000) 051a4f90ff14e2d6758a13397a92a69e taxon_horstiella_variabilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Horstiella variabilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Epicharis (Parepicharis) metatarsalis Friese, 1899 (type host) (Costa Rica); Epicharis (Parepicharis) zonata Smith, 1854 (Guyana, Bolivia); after Ochoa & OConnor, 2000 0b028d3e4428aa68996be4f0d38b87b0 taxon_horstiella_variabilis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Horstiella variabilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Often co-occurs with Horstiella concentrica on the same host, but topologically segregated from it. 5fe876525c7c69dc4f2c218388746562 taxon_laelapidae http://purl.org/dc/dcmitype/StillImage image/jpeg Pneumolaelaps costai, female, ex Bombus fervidus, USA: Ohio, BMOC 02-0706-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pneumolaelaps_costai_BMOC_02-0706-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3d6ee874c10847e0695f58615f441e2d taxon_laelapidae http://purl.org/dc/dcmitype/StillImage image/jpeg Eumellitiphis mellitus, male, ex Lestrimelitta limao, Panama, BMOC 88-0729-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Eumellitiphis_mellitus_BMOC_88-0729-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8d68ddc47bcd79e5241c18a20d238cc6 taxon_laelapidae http://purl.org/dc/dcmitype/StillImage image/jpeg Laelaspoides sp., phoretic female, ex Caenaugochlora costaricensis, Costa Rica, BMOC 80-0723-003 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Laelaspoides_sp_BMOC_80-0723-003.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 50d8ceafe181a82478cfc48ff348b07c taxon_laelapidae http://purl.org/dc/dcmitype/StillImage image/jpeg Urozercon sp, phoretic deutonymph, ex Scaura latitarsis, Costa Rica, BMOC 90-0101-020 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Urozercon_sp_BMOC_90-0101-020.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 5cef7c96afe20a3e568c40af71130845 taxon_laelapidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Laelapidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family is the most morphologically and ecologically diverse group of Mesostigmata. Thirty-five genera of laelapids have developed ectoparasitic associations with mammals; 10 genera are free-living predators in soil; 43 genera include species associated with arthropods: Coleoptera (8 mite genera), Blattaria (2), Isoptera (1), Hymenoptera (21), Araneae (1), Myriapoda (7) and terrestrial Crustacea (2) (Casanueva, 1993). Ecology of bee-associated mites is poorly known. Mites of the genus Neohypoaspis are predators on astigmatid mites in nests of stingless bees (Delfinado-Baker et al., 1983). Species of the genus Pneumolaelaps feed on pollen and nectar in nests of bumblebees (Royce and Krantz, 1989). Melittiphis alvearius is a scavenger on pollen in colonies of the European honeybee (Gibbins and Toor, 1990). Mites of the genera Varroa, Euvarroa,Tropilaelaps are obligatory parasites of different species of Apis, inflicting significant damage to apiculture (Eickwort, 1988). 22f172d09bc401eb7e8414ded584a554 taxon_laelaspoides http://purl.org/dc/dcmitype/StillImage image/jpeg Laelaspoides sp, phoretic deutonymph, ex Caenaugochlora costaricensis, Costa Rica, BMOC 80-0723-003 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Laelaspoides_sp_BMOC_80-0723-003.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e fcb190b77625d7ae2c7248a07016e559 taxon_laelaspoides http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Laelaspoides http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Monotypic genus restricted to augochlorine bees (Halictidae: Augochlorini) in North America. Pollenophagous. d48d23a08ce4c9ac47fbee7a4c5a8c40 taxon_laelaspoides_ordwayae http://purl.org/dc/dcmitype/StillImage image/jpeg Laelaspoides sp, female, ex Caenaugochlora costaricensis, Costa Rica, BMOC 80-0723-003 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Laelaspoides_sp_BMOC_80-0723-003.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b0c7a5d0254fb316c8c00ec5bb7e937b taxon_laelaspoides_ordwayae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Laelaspoides ordwayae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Kansas 8f829851f511ff6fe5640a3b2c54b4d1 taxon_laelaspoides_ordwayae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Laelaspoides ordwayae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Augochlorella striata (Provancher, 1888)
Augochlorella persimilis (Viereck, 1910)
9a9fcb8be2c11275423ccc2ebf81c9d6 taxon_laelaspoides_ordwayae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Laelaspoides ordwayae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology To date, Laelaspoides ordwayae is known in associations with two ground-nesting bee species, Augochlorella striata (Provancher, 1888) and Augochlorella persimilis (Viereck, 1910) in Kansas. The mites in all stages, except for larvae, are abundant in the nest cells of the bees and are encountered more rarely in the cell cluster cavity of the nests. Probably the larval period is short and the larvae moult into protonymphs soon after hatching, or that the mite is nymphiparous, passing the larval stage within the female. After the hibernation, and before the bees start to build nests, the mites disperse on bees via phoresy, holding primarily onto the hairs of the host. The mites probably enter the nest cells as they are being provisioned, reproduce there, then leave the cells as the young emerging bees open them, and enter other cells as they are provisioned. The mites eat pollen and apparently do not harm the bee larvae (Eickwort, 1966). 485300ba717f30d2bb6988d4825ac823 taxon_lasioseius http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Lasioseius http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Lasioseius, with approximately 100 described species, is probably the most morphologically diverse genus of Ascidae (Halliday et al., 1998). Species of this genus were found on rotting organic substances, in forest litter, moss, soil, in nests of small mammals and birds, under bark, in bracket-fungi, tree-holes, stored products, in warehouses, and hay. Several species were collected in beehives: Lasioseius penicilliger Berlese, 1916 in Canada, the former Czechoslovakia, and Iran (Crozier, 1989; Haragsim et al., 1987; Mosaddegh, 1997); Lasioseius berlesei (Oudemans, 1938) in Europe and Canada (Homann, 1933; Walter & Lindquist, 1989), and Lasioseius furcisetus Athias-Henriot, 1959 in the Western part of the former USSR (Bregetova, 1977). Lasioseius sp. was recorded from the Philippines in association with Apis cerana and Apis mellifera (Malabanan and Corpuz-Raros, 1998). There is no evidence, however, that beehives are their primary habitat. 5f2f7798e1cc56dda7f14333b600492c taxon_lasioseius_penicilliger http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Lasioseius penicilliger http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Nova Scotia; Mexico; British Isles; Italy (type locality); Germany; former Czechosovakia, Iran, New Zealand (Bregetova, 1977; Crozier, 1989; Estebanes-Gonzalez, 1997; Haragsim et al., 1987; Hughes, 1976; Kamali at al., 2001; Macfarlane, 2005; Mosaddegh, 1997). e270c755ca7fd34fc317ac3e8ee1b0dc taxon_lasioseius_penicilliger http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Lasioseius penicilliger http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species was found in decaying wood, old grassland and fields, in soil, nests of mammals and birds, in mills, on damp barley, and in beehives (Apis mellifera) and bumble bee (Bombus) nests. No harm to the bees was reported. Hughes (1976) speculates that this species might feed on mould. e5190f19b4f77fb08da899179aaa4b58 taxon_leptus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Leptus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Vercammen-Grandjean (1973) wrongly cites the year of the original proposal of Leptus as 1756. 56473229c0d2cd8dfecb13d4ffbf2d33 taxon_leptus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Leptus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Larvae of Leptus are known to parasitize different terrestrial arthropods. Two described species of this large genus occur on bees: Leptus ariel Southcott, 1989 was found on the European honey bee in Guatemala. Leptus monteithi Southcott, 1993 was described from a colletid bee Leioproctus sp. (Hymenoptera: Colletidae) from Australia (Tasmania) (Southcott, 1993) . Unidentified species were reported from the European honeybee in Peru (Cerro de Pasco) and Colombia (Fletchtmann, 1980; Losada, 1947; Southcott, 1989; 1993), and the USA (Wilson et al., 1990). a74f86a75ba69f4f03696bf7e8eb8cbe taxon_leptus_ariel http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Leptus ariel http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Guatemala (type locality); Peru (Haitlinger, 1999; Southcott, 1989). 869942c7ebecc6318782de66b45baac2 taxon_leptus_ariel http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Leptus ariel http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758 (Hymenoptera: Apidae) (type host)
Eumolpinae (Coleoptera: Chrysomelidae) (Haitlinger, 1999; Southcott, 1989).
66261119f4cccbf5a409871543b38791 taxon_leptus_ariel http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Leptus ariel http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Larvae were found attached to the hind femur, front head, and first metasomal tergite of the European honeybee Apis mellifera L. (Hymenoptera: Apidae). Since no adult mites were seen in the bee brood, it was assumed that mite propagation occurs outside the hive. The larval mites may have gained initial contact with the host while the bees were foraging on floral parts or while standing on moist soil investigating water. Observations within the hive did not reveal any obvious damage to the adult bees (Southcott, 1989; Wilson et al., 1987). Haltinger (1999) reported this species on a chrysomelid beetle in Peru.
4182122be0a481a10dcbd7222c5e0d46 taxon_leptus_monteithi http://purl.org/dc/dcmitype/StillImage image/jpeg Leptus monteithi, larva, ex Leioproctus sp. (Colletidae), Tasmania (modified from Southcott, 1993) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Leptus_monteithi.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 696d2be3c823b311b70c63c52eb8c1de taxon_leptus_monteithi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Leptus monteithi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia: Tasmania, Mt Barrow, nr Launceston (type locality) 75d06d7e20a320a76692e0c1a8c89314 taxon_leptus_monteithi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Leptus monteithi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Leioproctus sp. (as Lepioproctis (Leioproctis) sp.) a3d021b597abe8f224a252d336093168 taxon_locustacarus http://purl.org/dc/dcmitype/StillImage image/jpeg Locustacarus buchneri (Podapolipidae), larvifrom female, dorsal view title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Locustacarus_buchneri.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f87ba5c97f8900fecfc78567864288fe taxon_locustacarus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Locustacarus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus includes three species. Locustacarus trachealis and L. masoni Husband, 1974 occur in the respiratory system of their orthopteran hosts apparently piercing the tracheal walls to feed. Locustacarus buchneri is an internal parasite of bumblebees (Husband, 1974; Husband and Sinha, 1970). f4b33c477fed5009ac3f54e7f3fac287 taxon_locustacarus_buchneri http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Locustacarus buchneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The type locality and host are uncertain because the holotype has not been designated and multiply localities and hosts have been reported in the original description. Localities (original order): Franken, Mecklenburg (Germany), Tirol (France). Hosts (original order): Bombus pratorum, B. barbutellus (in Psithyrus), B. sylvarum, B. pascuorum (as agrorum), B. hortorum, B. lapidarius, B. terrestris, B. lucorum, B. ruderarius (as derhamellus), B. veteranus (as equestris), B. campestris (in Psithyrus), B. vestalis (in Psithyrus). 4dbe3c403964da50d89e441a70e58bc1 taxon_locustacarus_buchneri http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Locustacarus buchneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Manitoba; USA: Michigan, Colorado, Missouri, Virginia, Kansas; England; Belgium; Netherlands; France, Germany; former Czechoslovakia; Denmark; Sweden; Poland; Russia; Armenia; Japan; India; New Zealand (Eidelberg, 1994; Grobov, 1975; Goldblatt & Fell, 1984; Goka et al., 2001, 2002; Husband & Husband, 1996; Husband and Sinha, 1970; Macfarlane, 1975). 19446a772711ef5c1263cb50c7ef943a taxon_locustacarus_buchneri http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Locustacarus buchneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Holarctic:; Bombus (Alpinobombus) balteatus Dahlbom, 1832; Bombus (Bombus) terricola Kirby, 1837; Bombus (Psithyrus) citrinus (Smith, 1854), reported as "Psythyrus laboriosus F."; Bombus (Pyrobombus) bimaculatus Cresson, 1863; Bombus (Pyrobombus) perplexus Cresson, 1863; Bombus (Pyrobombus) vagans Smith, 1854; ; Palaearctic:; Apis mellifera Linnaeus, 1758 (probably accidental record); Bombus (Alpinobombus) balteatus Dahlbom, 1832; Bombus (Alpinobombus) polaris Curtis, 1835; Bombus (Bombus) hypocrita Pérez, 1905; Bombus (Bombus) ignitus Smith, 1869; Bombus (Bombus) lucorum (Linnaeus, 1761); Bombus (Bombus) terrestris (Linnaeus, 1758); Bombus (Bombus) tunicatus Smith, 1852; Bombus (Megabombus) consobrinus Dahlbom, 1832; Bombus (Megabombus) hortorum (Linnaeus, 1761); Bombus (Melanobombus) lapidarius (Linnaeus, 1758); Bombus (Psithyrus) barbutellus (Kirby, 1802); Bombus (Psithyrus) campestris (Panzer, 1801); Bombus (Psithyrus) vestalis (Geoffroy, 1785); Bombus (Pyrobombus) hypnorum (Linnaeus, 1758); Bombus (Pyrobombus) pratorum (Linnaeus, 1761); Bombus (Thoracobombus) pascuorum (Scopoli, 1763) reported as "B. agrorum (Fabricius, 1787)"; Bombus (Thoracobombus) ruderarius (Müller, 1776) reported as "B. derhamelus K."; Bombus (Thoracobombus) sylvarum (Linnaeus, 1761); Bombus (Thoracobombus) veteranus (Fabricius, 1793) reported as "Bombus equestris F."; ; New Zealand (introduced hosts):; Bombus (Bombus) terrestris (Linnaeus, 1758); Bombus (Megabombus) ruderatus (Fabricius, 1775); ; (Husband & Husband, 1996; Husband and Sinha, 1970; Goka et al., 2001; Goldblatt & Fell, 1984; Macfarlane, 1975; Stammer, 1951; Tomaszewska, 1988) c8fe682e6ff6fd83558db16c5eb10646 taxon_locustacarus_buchneri http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Locustacarus buchneri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Locustacarus buchneri is an internal parasite of bumblebees occurring in tracheae and air sacs. Husband and Sinha (1970) reconstructed its life cycle. In the late April, the queen of Bombus bimaculatus emerges from overwintering sites in the soil. Within a few days, female mites in the tracheae enlarge as the haemolymph is ingested and ovaries begin to function. Several female mites may be found in tracheae of each queen bee. Nearly one week after emergence of a queen bee, eggs hatch. As many as 50 eggs were associated with a single female mite. Paired male and larviform female mites may be found among the eggs. In some cases the sex ratio of young mites is near unity. Some females may have been unfertilized and may produce only males. Males have poorly developed mouthparts, are short lived, and do not leave the host bee. Larviform female mites migrate to the tracheae of worker bees, attach to the tracheal walls, molt, and begin to enlarge. Adult females with only two legs and a greatly distended body cannot leave the host. This cycle may be repeated several times during the next eight weeks. In the first week in July, male bees and new queen bees emerge. Males of Psithyrus spp., B. vagans, and other B. bimaculatus occasionally enter the nest and may bring in or take out larviform female mites in the tracheae. By early August, new queen bees have mated, left the nest, and some may have entered diapause at sites several inches below the ground. After the new queen bees leave the nest, few new workers are produced and older workers accumulate more and more mites. The numbers of mites in worker and male bees in August is sufficient to severely injure the bees. Diarrhea is observed and some bees are lethargic and no longer forage. It was at this time that a small male mite was found. This observation is slight evidence that small males may be produced by crowding of mites, poor nutrition, or by aging of the parent. Because worker bees, male bees, and the old queen bee die during fall, the only mites which survive to the following April are in the tracheae of young queens in diapause.
Goka et al., 2001 reported introduction of this species to Japan with commercially breeding Bombus terrestris from Belgium and Netherlands. Tomaszewska (1988) reported this species for the honey bee, Apis mellifera.
1c937dff37eb41ca50167414900651ea taxon_macrocheles http://purl.org/dc/dcmitype/StillImage image/jpeg Macrocheles praedafimetorum (Acari: Macrochelidae), female, ex nest of the bumblebee Bombus terricola,  Canada: Alberta title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Macrocheles_praedafimetorum.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 011934f5ce488fd9860bb19cf16f07ee taxon_macrocheles http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Macrocheles http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The type species of this genus is colloquially considered (e.g., Oudemans, 1936; Karntz, 1962; Bregetova, 1977; Evans and Till, 1979) as Acarus marginatus Hermann, 1804 (=Acarus muscae domesticae Scopoli, 1772) with the reference to Latreille in Cuvier & Latreille, 1829 (p. 282). However, in this work, two species are cited under Macrocheles Latr.: "Je rapporte à ce genre les Acarus marginatus et testudinarius d'Herman fils (Mémoire aptérol. pag 76, pl. VI fig 6, et pag 80 pl. IX fig. 1)" (p. 282). Because two species were originally placed in Macrocheles and none of them has been indicated as the type, the type designation for this genus should be attributed to the first reviewer. It is beyond the scope of our project to make a detailed investigation on this matter, but we note that Oudemans (1936, p. 169) indicated Drapiez (1841) who explicitly mentioned Acarus marginatus Hermann as the type (p. 19, "L'Acarus marginatus d'Hermann forme le type de ce genre"). e872cb7bdd847a3397e8a39c648e2351 taxon_macrocheles http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Macrocheles http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This large cosmopolitan genus includes species that inhabit a variety of habitats: soil, humus, forest litter, moss, dung, hay, rotting stumps, galleries of bark beetles and subcortical situations, nests of rodents, birds, mammals, and bumblebees. The mites are often phoretic on insects of the orders Coleoptera and Diptera. Two, apparently related, species are known from bumblebees: Macrocheles rotundiscutis Bregetova and Koroleva, 1960 (=Macrocheles bombophilus Goetz, in Hirschmann, 1970) from Europe and Macrocheles praedafimetorum Richards and Richards, 1977 from North America (see biological account on this species here). Three other species, Macrocheles muscaedomesticae (Scopoli, 1772), Macrocheles nataliae Bregetova et Koroleva, 1960, Macrocheles glaber (Müller, 1860), and Macrocheles martius (Hull, 1925), have been reported from the honey bee, Apis mellifera L. (Banaszak, 1980; Chmielewski, 1991; Haragsim et al., 1978). The two latter species are incidental inhabitants of hives, and they probably use honeybees for transport (Chmielewski, 1991). Lo and Chao (1975) reported a macrochelid mite (probably a Macrocheles) in Taiwan that occurs in the flowers of loquat (Eriobotrya japonica Lindl. (Thunb.) (Rosales: Rosaceae)) in the spring and uses both indigenous (Apis cerana Fabricius, 1793) and introduced (Apis mellifera) bee species for dispersal. The following list summarizes records of Macrocheles from bees and their nests: 9fe6122e5b1d1d4a2329bed980afd706 taxon_macrocheles_praedafimetorum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Macrocheles praedafimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Alberta. 9ceec7c4a55b87370b0fd7e4790ca50e taxon_macrocheles_praedafimetorum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Macrocheles praedafimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758; Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Pyrobombus) bifarius Cresson, 1878; Bombus (Pyrobombus) centralis Cresson, 1864; Bombus (Pyrobombus) flavifrons Cresson, 1863 (type host); Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) melanopygus Nylander, 1848; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878
c6bf043c7929c9e714200f6d1017e57a taxon_macrocheles_praedafimetorum http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Macrocheles praedafimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Macrocheles praedafimetorum is a polyxenous species occurring in the nests of many species of bumble bees, although fewer in the nests of Bombus rufocinctus and B. appositus, which emerge and construct nests later in the spring.
The mated female mite is the phoretic stage. Bumble bees are not the vector on which this mite entered the nests. A dung-inhabiting beetle, Onthophagus nuchicornis L. (Coleoptera: Scarabaeidae), was observed entering a bumble bee nest with three female mites attached. This beetle is apparently attracted to the bumble bee nests by the decaying material in the bottom of the nest.
Macrocheles praedafimetorum is a predatory mite. It can feed on the nematode Caenorhabditis sp., which occurs in profuse numbers in moist rotting material in bumble bee nests. The mite was also observed feeding on the eggs, larvae, and adults of Coboldia fuscipes (Meigen) (Diptera: Scatopsidae) and an unidentified psychodid. Larvae of these flies feed on the decaying nest material, and populations are high in some nests. These mites are cannibalistic; newly molted protonymphs were observed eating larvae, while females ate larvae and protonymphs. Perhaps in response to this behavior, females laid up to 10 eggs inside empty fly pupal cases.
Macrocheles praedafimetorum is an arrhenotokous species, the unfertilized females producing only males and fertilized females giving rise to both sexes. Average life span for males is 21.8 days (range 6-49, n=19), which is significantly shorter than that for females, 36.2 days (range 13-73, n=9). Females more commonly occur in the nests, because the females live longer in the phoretic stage and withstand greater environmental fluctuations (i.e., reduced humidity and food) than the males. Most females must mate before dispersing as they produce both males and females upon arrival at the new food source. Males apparently sense when female deutonymphs are about to molt. The male climbs on the female's back and remains until she molts, at which time it crawls under the female, positioning itself so that its chelicera meet the female's genital plates. Spermatophores are then transferred by the chelicera.
The mites are also common in artificial domiciles. Underground domiciles were preferred probably because the early nesting bumble bee species that occupied these domiciles had larger populations resulting in more accumulating debris and decaying material. The mite does not appear to prefer domiciles placed in open meadows or above ground. Those domiciles were usually drier, had less decaying material, and fewer debris-feeding organisms. The mite was also found in cattle dung.
Macrocheles praedafimetorum has no direct effect on the bumble bee population within the nest. The mite exploits a wide range of saprophagous organisms common to dung, decaying matter, bumblebee and honeybee nests (Richards and Richards, 1977; Crozier, 1989). f036a24e413d1c47cace523be6452ba0 taxon_macrochelidae http://purl.org/dc/dcmitype/StillImage image/jpeg Macrocheles praedafimetorum (Acari: Macrochelidae), female, ex nest of the bumblebee Bombus terricola,  Canada: Alberta title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Macrocheles_praedafimetorum.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 95e10560a33c0a447ab34b646aa5e784 taxon_macrochelidae http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis sp., ex nest of the stingless bee Lestrimelitta limao, Panama, BMOC 88-0729-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_sp_BMOC_88-0729-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 60321b17805f361ea53cb6fb373a509c taxon_macrochelidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Macrochelidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes 14 genera. Two of them, Trigonholaspis and Macrocheles, were recorded from bees (Chmielewski, 1991; De Jong et al., 1982; Eickwort, 1988; Krantz, 1998; Richards and Richards, 1977). 1b3c9726420cdebceadd2c2e412f7ea5 taxon_macrochelidae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Family Macrochelidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution

Key to genera of Macrochelidae associated with bees

1. Four pairs of opisthonotal dorsocentral setae (J1, J2, J3, J5). Anterodorsal "hump" present. Sternal shield not coarsely punctate ... Trigonholaspis

- Two, rarely three, pairs of opisthonotal dorsocentral setae. Anterodorsal "hump" absent. Some areas of sternal shield often coarsely punctate... Macrocheles 76cb67906f21ee242d5e40f4ac1a9db8 taxon_melissotydeus http://purl.org/dc/dcmitype/StillImage image/jpeg Melissotydeus macrosolenus, female, dorsal view, ex Partamona mourei, Brazil (modified from André, 1985) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Melissotydeus_macrosolenus_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e cc9927a6a353cedc01eb7566289f299d taxon_melissotydeus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Melissotydeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus includes two species. Melissotydeus macrosolenus André was collected from nests of Partamona and Trigona (Apidae, Meliponini) in Brazil. Melissotydeus incarum Kazmierski, 1998 was found among debris in dense thickets of bamboo in Peru (André, 1985; Kazmierski, 1998a). 80f63c1cc4d2e877847c5cb7e2cf92b9 taxon_melissotydeus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Melissotydeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil, Peru (André, 1985; Kazmierski, 1998a). f9a2e56d82c66e6746fe252ddd1a145a taxon_melissotydeus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Melissotydeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Partamona mourei Camargo, 1980 (type host); Frieseomellita sp. (as "Frieseomellita modesta" (nom. nud., Moure’s ms n., see Moure at al., 2011]) (André, 1985). f08ba26246094b09819020a8ed7f43db taxon_melissotydeus_macrosolenus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Melissotydeus macrosolenus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Roraima: Mucajai (type locality), Amazonas (André, 1985). 60ae632f85be12815806896c747da965 taxon_melissotydeus_macrosolenus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Melissotydeus macrosolenus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Partamona mourei Camargo, 1980 (type host)
; Frieseomellita sp. (as “Frieseomellita modesta" (nom. nud., Moure’s ms n., see Moure at al., 2011]) (André, 1985).;
efd02e8c6cebfa8da66135bd4b9b740d taxon_melissotydeus_macrosolenus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Melissotydeus macrosolenus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Eickwort (1994) hypothesized that this species may be mutualistic feeding on fungi in the bee's nests. a3141bff1d8cbd8312f8f144e26cc0ca taxon_melittiphis http://purl.org/dc/dcmitype/StillImage image/jpeg Mite Melittiphis alvearius (Laelapidae) found in beehives. Modified from Evans and Till (1966) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Melittiphis_alvearius.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 0f93ddd630567fb0057c57bb56fbffad taxon_melittiphis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Melittiphis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Monotypic genus. Pollenophagous in beehives of the European honey bee, Apis mellifera. be141ea5241c9457e4816b8b168e5266 taxon_melittiphis_alvearius http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Melittiphis alvearius http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Nova Scotia, Ontario; Chile; Italy (type locality), England; Germany; former Czechoslovakia; Russia; Australia; New Zeland (Berlese, 1918; Casanueva, 1992; Crozier, 1989; Davis and MacRory, 1987; Delfinado-Baker, 1994; Evans and Till, 1966; Gibbins and Toor, 1990; Grobov, 1975; Knihinicki and Halliday, 1995; Samšiňák et al., 1978; Vitzthum, 1930). 01fe4235368c58f0e4a91f98dfe34093 taxon_melittiphis_alvearius http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Melittiphis alvearius http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758 (type host)
e46c2ad9d05bf7a2998d6820b4fe272f taxon_melittiphis_alvearius http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Melittiphis alvearius http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in beehives of the European honey bee, Apis mellifera or on bees. Serological procedures demonstrated that this mite is not a parasite or predator, but rather a scavenger of pollen in the hives (Gibbins and Toor, 1990). 091cd09c53b79510c4b993a76ef3cea1 taxon_neohypoaspis http://purl.org/dc/dcmitype/StillImage image/jpeg Mite Neohypoaspis ampliseta (Laelapidae) associated with Trigona spp. (Apidae: Meliponini) in Panama. Modified from Delfinado-Baker et al., 1983 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Neohypoaspis_ampliseta.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 34b8d1bac045ed6609eeafbedfa9bb8c taxon_neohypoaspis http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Neohypoaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Monotypic genus. Associated with obligate necrophagous Trigona spp. in the Neotropics. Probably beneficial, feeding on astigmatid mites that consume honey in the nest (Delfinado-Baker et al. 1983). df3079f77e61edb1b96f3d5ac399529c taxon_neohypoaspis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Neohypoaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Neotropics dd6d0b5abb9086a26d2b51bb745973bc taxon_neohypoaspis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Neohypoaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona spp. b086f52b0241d375bb2c4cab8156fbc6 taxon_neohypoaspis_ampliseta http://purl.org/dc/dcmitype/StillImage image/jpeg  Mite Neohypoaspis ampliseta (Laelapidae) associated with Trigona spp. (Apidae: Meliponini) in Panama. Modified from Delfinado-Baker et al., 1983 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Neohypoaspis_ampliseta.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f8481dbcafa91bb4ff837b4664cb848d taxon_neohypoaspis_ampliseta http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Neohypoaspis ampliseta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Panama: Panama Prov. (type locality). 8bcb29c191303a5e922776cb739950f9 taxon_neohypoaspis_ampliseta http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Neohypoaspis ampliseta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Trigona fulviventris Guérin-Méneville, 1844
Trigona hypogea Silvestri, 1902 (type species)
345100af809666964b3030c3856dc9fb taxon_neohypoaspis_ampliseta http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Neohypoaspis ampliseta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species occurs in great numbers in the outer sheath of the nests of two species of stingless bees, Trigona fulviventris and Trigona hypogea. It is worth noting that the last bee species is obligate necrophage (Camargo, 1991). The mites were located in the base of the nests, often in areas that can not be reached by the bees. It was shown in the laboratory, that these mites are predators on astigmatid mites normally found in the nests and do not consume pollen or honey. Thus, Neohypoaspis ampliseta is probably beneficial for its bee hosts (Delfinado-Baker et al., 1983).
ea69d17d121aacbdb198bc4cc2c7a26e taxon_neotrombidiidae http://purl.org/dc/dcmitype/StillImage image/jpeg Prodorsal sclerite, eyes, and anterior hysterosomal setae of the mite Neotrombidium sp. (Neotrombidiidae), larva, ex Xylocopa micans, USA: Florida, BMOC 03-0525-002 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Neotrombidium_sp_larva_BMOC_03-0525-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a647472bc507c26ca483bc411fb3486b taxon_neotrombidiidae http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Family Neotrombidiidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Palpal genu with one seta. One pair of dorsal trichobotria on prodorsal sclerite. Scutellum bearing setae c1 absent. Claparede's organ present. Postero-apical corner of at least posterior coxa with distinct reticulate pattern. Femur I with at least one solenidia and seven pilose setae. Femora II-III with at least two solenidia. Only one empodial claw present on each tarsus.
Notes. Some authors (e.g., Zhang et al., 2011) cite the year for this family as 1959. However, this family name was first proposed in 1955 (Feider, 1955). 68df69568b6bbcbbf6ebb89d6527ed91 taxon_neotrombidiidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Neotrombidiidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes five genera and 24 species (Feider, 1977; Krantz, 1978; Vercammen-Grandjean, 1973; Zhang et al., 2011). Larvae are parasitic on Coleoptera, Diptera, and rarely on Hymenoptera (our data), while postlarval stages, including adults, live in forest litter and under bark of trees where they presumably prey on other small arthropods. The family is worldwide in distribution, excluding boreal and polar regions (Lindquist and Vercammen-Grandjean, 1971; Singer, 1971). 93e45dfce7b264cc3001b61bae282711 taxon_neotrombidiidae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Family Neotrombidiidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa micans Lepeletier, 1841. c8f3a67a38bc5d6a8a1121e8d2b61fa1 taxon_neotrombidium http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Neotrombidium http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description The femur is subdivided onto telo- and basifemur. See Lindquist and Vercammen-Grandjean (1971) for other characters. 09f7b7a16d6df371291532196c5ed0aa taxon_neotrombidium http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Neotrombidium http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus is worldwide in distribution and includes 16 species (Beron, 1995; Lindquist and Vercammen-Grandjean, 1971; Singer, 1971). Larvae are parasitic on unrelated families of subcortical Coleoptera: Cerambicidae, Elateridae, and Cleridae (Lindquist and Vercammen-Grandjean, 1971). We found a single larva of this genus on the bee Xylocopa micans. The level of association of this mite species with the host, thus, is unknown. f325acc9fa3f2d06728057dcb37b9bbd taxon_neotrombidium http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Neotrombidium http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa micans Lepeletier, 1841. 92e4c63b4c51478b1ddaabb23f708784 taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Uropore absent. Terminal setae of palpotarsus not different from other setae of palpotarsus, pectinate. Anterior sternal setae not fused with coxal fields I. Two nude genualae on leg I. Femoralae III not reaching apex of genu. Genuala III almost reaching apex of tibia. f3229185dea1af0149c855efe0b145fb taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Florida. ce74576af9acd6d734a723b70de3239e taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa micans Lepeletier, 1841. d04ff12c1531decce1207d788ce7f436 taxon_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology A single larva was found attached to the soft membrane in the region of hind coxae of the bee host. 26b6a8a9a290396a57d037dbbbda64a0 taxon_nodele http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic females, on propodeum of Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_on_bee_e_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e4f5a78ce2f127a6e15ba02507d48703 taxon_nodele http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Nodele http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes five species collected from forest litter, under bark, on bostrichid beetles, galleries of bark beetles, nest of sparrow (Passer), and grain in the Holarctic, Oriental, and Afrotropic realms (Fain and Bochkov, 2001a, Kuznetsov, 1977; Thewke and Enns, 1968; Wafa and Soliman, 1968).
Biology is only known for Nodele calamondin. It is a predator of other mite species and of small insect larvae. Unmated females produced only male offspring (arrhenotoky), which subsequently mated with their mothers to produce female as well as male offspring. Like other representatives of the family, males have one nymphal stage in their development and females have two nymphal stages. Males guarded the quiescent deutonymph females and mating took place as soon as the females emerged as adults. Unmated females lived more than twice as long as mated females and laid about 89% of the number of eggs laid by their mated sisters. The average duration of a generation was 62 days at 25 degrees C and 34 days at 29.5 degrees C. Net reproduction rate (ratio of female births in two successive generations) was 20 at 25 degrees C and 34 and 29.5 degrees C (after Barker, 1992). There is a single record of three females of Nodele (Aztecocheyletus) conquistador collected from the propodeum of the megachilid bee Aztecanthidium tenochtitlanicum (Bochkov & Klimov, 2004). 38c988e4924ab9bdef2fdb89cf52e3d0 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e d67c9b264b666417bfbbe1061116e793 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female, dorsal view, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_d_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7ab22f0263b992567f041e666e2ed201 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female, ventral view, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_v_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 80dc5b86002863351551762cf1d3cc7d taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female on host, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_on_bee_b_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ee710ca0e9f2998c9c54931409b2b775 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female on host, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_on_bee_d_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a88a9370ea1032e3309070d27b6baab7 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/StillImage image/jpeg Nodele (Aztecocheyletus) conquistador, phoretic female on host, ex Aztecanthidium tenochtitlanicum, Mexico, BMOC 03-1003-043 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Nodele_conquistador_on_bee_e_BMOC_03-1003-043.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2438fedee9d0c65757a29fad9f65efb4 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Nodele (Aztecocheyletus) conquistador http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Two pairs of setae of series E, e1 and e2. All dorsal idiosomal setae clavate. Protegmen bears a pair of well-developed lateral teeth. Propodonotal shield transversely serrate-striated. Tibia I relatively short. Tarsus I, excluding pretarsus/tibia I 2.3 : 1.0. Solenidion w1 short, 4 times shorter than seta ft, situated far from guard seta. 94aead516b59ad71b8442d05482cb3cd taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Nodele (Aztecocheyletus) conquistador http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Jalisco. 60334a3fa27f3cb11b7cb6fc796cfac3 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Nodele (Aztecocheyletus) conquistador http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Aztecanthidium tenochtitlanicum Snelling, 1987. 54715cadcf7d813c6d1006ab86724fe2 taxon_nodele_aztecocheyletus_conquistador http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Nodele (Aztecocheyletus) conquistador http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology We believe that relationships of our species with the bee, like in other species of the genus Nodele, depend simply on availability of a host in the mite environment. In other words, it is not host specific. a8eaa80428ccfedbaf0d5522b2539283 taxon_parasitellus http://purl.org/dc/dcmitype/StillImage image/jpeg Parasitellus talparum (=Parasitus favus) (Acari: Parasitidae), phoretic deutonymph, ex Bombus flavifrons, USA: Wyoming, BMOC 99-0624-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Parasitellus_talparum_dn_BMOC_99-0624-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ef67ba934cba948d85c0e00678be1f6c taxon_parasitellus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Parasitellus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes 11 species that inhabit nests of bumblebees (Bombus) in the Holarctic region (Davydova, 1988; Hyatt, 1980; Karg, 1985; Richards, 1976). Occasionally they occur in beehives or burrows of small mammals (Hyatt, 1980; Crozier, 1989). Deutonymphs are commonly phoretic on the adult bumblebees or cuckoo bumblebees. Since bumblebee colonies are annual and only young queens overwinter, mite deutonymphs are able to distinguish between queens and other castes (workers, males). Although the mites may disperse on all castes of bumblebees, they prefer queens, and never move from a queen to a male. Mites dispersing on workers and males may try to switch to queens later, either during copulation or on flowers, where bumblebees forage (Huck et al., 1998; Schwarz and Huck, 1997).
The species of Parasitellus, although only associated with bumblebees, are not specific to a particular species of host, with species often co-occurring in individual Bombus nests. The lack of host specificity in this group may be the result of host switching in flowers by cockoo bumblebees, or by queens that visit old nests with overwintered deutonymphs. In the nests of a single bumblebee species, mites can disperse due to queen supersedure or invasion of workers or queens from different nests (Richards and Richards, 1976; Tichomirov, 1969a).
The exact nature of the association between these mites and their bumblebee hosts is uncertain, although predatory behavior toward acarid mites and other parasitid mites was suggested (Richards and Richards, 1976). If these mites feed preferentially on potentially damaging acarid mites, they may be beneficial to colony health. In Parasitellus fucorum, males, larvae, protonymphs, and possibly deutonymphs were found to be predatory under laboratory conditions, while pollen was the preferred food for adult females and deutonymphs (Koulianos and Schwarz, 1999).
Variation in many species of Parasitellus involves asymmetry in the shape of the tectum. 969e77fed72fd4eb48e1797643b8cb52 taxon_parasitellus_hobbsi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Parasitellus hobbsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Alberta (type locality); Nova Scotia (Crozier, 1989; Richards, 1976). 022f18075ab632cdd8d5382be9c599fa taxon_parasitellus_hobbsi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Parasitellus hobbsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758 Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Psithyrus) fernaldae (Franklin, 1911); Bombus (Psithyrus) insularis (Smith, 1861); Bombus (Pyrobombus) bifarius Cresson, 1878 (type host); Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878; (Richards, 1976; Rrichards and Richards, 1976; Crozier, 1989)
5966912b3b512ea2c38010e6ab26138b taxon_parasitellus_hobbsi http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Parasitellus hobbsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Similar to that of other Parasitellus. Occurs less frequently than P. talparum and P. perthecatus. In Canada (Nova Scotia) was found in beehives. 94f3a6dd6f101c4af24ed03f0697d7ae taxon_parasitellus_inquilinobombus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Parasitellus inquilinobombus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA (first record): Michigan; Canada: Alberta (type locality). a15aba31aee219a87bd949fefa0c53e6 taxon_parasitellus_inquilinobombus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Parasitellus inquilinobombus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Psithyrus) fernaldae (Franklin, 1911); Bombus (Psithyrus) insularis (Smith, 1861); Bombus (Pyrobombus) bifarius Cresson, 1878 (type host); Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878; (Richards, 1976; Rrichards and Richards, 1976)
63d537c0b50ccbb8a8d0a5e74fff3afd taxon_parasitellus_inquilinobombus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Parasitellus inquilinobombus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Similar to that of other Parasitellus. Occurs less frequently than P. talparum and P. perthecatus. c7fede5293a15081d7511bb54c8c9143 taxon_parasitellus_perthecatus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Parasitellus perthecatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA (first record): Alaska, Ohio, Colorado; Canada: Northwest Territories, Alberta (type locality). 19f4eba8df636006728ece655aade7fe taxon_parasitellus_perthecatus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Parasitellus perthecatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Alpinobombus) polaris Curtis, 1835; Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Psithyrus) fernaldae (Franklin, 1911); Bombus (Psithyrus) insularis (Smith, 1861); Bombus (Psithyrus) suckleyi Green, 1860; Bombus (Pyrobombus) bifarius Cresson, 1878; Bombus (Pyrobombus) centralis Cresson,1864; Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) lapponicus (Fabricius, 1793) (as sylvicola); Bombus (Pyrobombus) melanopygus Nylander, 1848; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878 (type host); (Richards, 1976; Rrichards and Richards, 1976)
22806aea20a3ca6bef081052e5f9ea43 taxon_parasitellus_perthecatus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Parasitellus perthecatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Female lays 0-8 eggs per day in laboratory settings. Larvae hatch in 3-6 days (average 3.9) and in 3-6 days molt to protonymphs. The highest number of the deutonymphs on a single bee queen can be up to 210. Otherwise the biology is similar to that of P. talparum (Richards and Richards, 1976). 3d22fa0d48acb40be0b5c2ff2fc351c4 taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Michoacán de Ocampo. 8e6944d6542735b6f77f80193950ac07 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Bombus (Fervidobombus) diligens Smith, 1861. 736fc166c93ad47c39f389f8e03373a9 taxon_parasitellus_talparum http://purl.org/dc/dcmitype/StillImage image/jpeg Parasitellus talparum (=Parasitus favus) (Acari: Parasitidae), phoretic deutonymph, ex Bombus flavifrons, USA: Wyoming, BMOC 99-0624-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Parasitellus_talparum_dn_BMOC_99-0624-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 83e80d198f92c4e8bd4cfee89fa5696a taxon_parasitellus_talparum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Parasitellus talparum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA (first record): Alaska, Wyoming, Ohio; Canada: Alberta, Northwest Territories; Greenland; British Isles; Sweden; Germany; Netherlands (type locality); Switzerland; former Czechoslovakia; Russia: Moskovskaya Oblast', Western and Eastern Siberia, Kuril Islands; Turkmenistan; China: Qinghai Prov. (Hyatt, 1980; Gu et al., 1987; Klimov, 1998; Micherdzinski, 1969; Richards, 1976; Richards and Richards, 1976; Tichomirov, 1977). c278b88ef0d8933dea593894400d356c taxon_parasitellus_talparum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Parasitellus talparum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
North America:; Bombus (Alpinobombus) hyperboreus Schönherr, 1809; Bombus (Alpinobombus) polaris Curtis, 1835; Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Psithyrus) fernaldae (Franklin, 1911); Bombus (Psithyrus) insularis (Smith, 1861); Bombus (Psithyrus) suckleyi Green, 1860; Bombus (Pyrobombus) bifarius Cresson, 1878; Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) lapponicus (Fabricius, 1793) (as sylvicola); Bombus (Pyrobombus) melanopygus Nylander, 1848; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878; (Karg, 1985; Richards, 1976; Richards and Richards, 1976); ; Eurasia (only bumblebee hosts are listed):; Bombus (Bombus) albocinctus Smith; Bombus (Bombus) florilegus Panfilov, 1956; Bombus (Bombus) terrestris (Linnaeus, 1758); Bombus (Megabombus) tichenkoi (Skorikov, 1925); Bombus (Melanobombus) lapidarius (Linnaeus, 1758); Bombus (Thoracobombus) pascuorum (Scopoli, 1763) (as agrorum (F.)); (Hyatt, 1980; Klimov, 1998; Micherdzinski, 1969)
745495c4cacd0bc13c0a0ce27d2863de taxon_parasitellus_talparum http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Parasitellus talparum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species was originally described from a nest of the common mole, Talpa europea L., 1758 in the Netherlands. Numerous subsequent records indicate that this species is actually associated with different species of Bombus (Hyatt, 1980). The biology of Parsitellus talparum (=Parasitus favus) was studied by Richards and Richards (1976). In a bee nest, several generations of mites can be accomplished in a single season. Males copulate with females several times. Female lays 0-1 eggs in laboratory settings. Larvae hatch in 5 days and molt into protonymphs in six days. Protonymphs constantly die in cultures. The deutonymphs captured from bumblebee nests were able to survive longer in culture than any other developmental stages. Usually the sex ratio is biased toward females. The feeding preferences are not known, although feeding on nidicolous microarthropods is possible. Deutonymphs and females were observed to be cannibalistic. Deutonymphs usually attach themselves to hairs of metasoma and propodeum using their chelicerae. If their numbers are great, they will also crowd around the neck. Usually the mites occur on queens before the bee starts to rearrange nesting material in a prospective nest, and more rarely on workers or males. The number of deutonymphs increases with queen production. The highest number of Parasitus deutonymphs on a single bee queen can be up to 210, but usually they are less numerous. The mites are easily excited, and if disturbed will run over the entire body, but seldom leave the bee. 51891c3a9e77779c496ea61fc1567cc5 taxon_parasitidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Parasitidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family is distributed worldwide and includes 426 species and 35 genera grouped in 2 subfamiles: Parasitinae and Pergamasinae (Beaulieu at al., 2011; Johnston, 1982; Tichomirov, 1977). Parasitid mites are essentially predatory and feed upon other microarthropods, including their eggs, and on nematodes. They live in moss, forest litter, soil, dung, rotting seaweed, decaying organic substances, caves, and nests of small mammals and insects. These mites disperse during the deutonymph stage, usually on insects of the orders Coleoptera and Hymenoptera. All species of the genus Parasitellus are obligatory associates of bumblebees (Bombus) in the Holarctic region. Several occasional records of different species of the genus Parasitus and Vulgarogamasus are known from bumblebee nests and honeybee hives in North America and Europe. Pergamasus crassipes (Linnaeus, 1758), Pergamasus diversus Halbert, 1915, Poecilochirus necrophori Vitzthum, 1930, Holoparasitus sp., and Eugamasus sp., were recorded from the honeybee, Apis mellifera L. in Europe and Iran (Chmielewski W. 1991; De Jong et al., 1982; Haragsim et al., 1987; Grobov, 1975, 1978; Kamali at al., 2001; Mosaddegh, 1997), but these records probably are also accidental. 9ccb5fa4eeae8cb0ee64e3b89f94a0ec taxon_parasitus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Parasitus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Species of this genus live in manure, dung, compost or other decaying organic substances. Many of them have broad geographic ranges. Usually, deutonymphs are phoretic on dung beetles (Hyatt, 1980).
Several occasional records are available from bumblebee nests: Parasitus fimetorum (Berlese, 1904) in North America and Europe (Hyatt, 1980; Richards, 1976); Parasitus beta Oudemans et Voigts, 1904, Parasitus coleoptratorum (Linnaeus, 1758), Parasitus cornutosimilis (Schweizer, 1949), Parasitus consanguineus (Oudemans & Voigts, 1904) and Parasitus loricatus (Wankel, 1861) in Europe (Hyatt, 1980; Karg, 1971). Parasitus coleoptratorum and Parasitus distinctus Berlese, 1903 were also found in honeybee hives (Apis mellifera) in Europe (De Jong et al., 1982; Grobov, 1975), and we have seen a single female of Parasitus aff. loricatus from a mammal nest in a bee hive (New York, Tompkins Co., Ithaca, 14 Dec 1976, V. Lazar, BMOC 76-1214-001).
We give an account on Parasitus fimetorum, because it frequently occurs in the nests of bumblebees in the United States and Canada (Richards, 1976; Richards and Richards, 1976). Athias-Henriot (1979) created a new genus for this species, Phorytocarpais Athias-Henriot, 1979, however, we accept here generic designation of Hyatt (1980). 2c1d4c78e5a5e6d09e898ebcf6c1679c taxon_parasitus_fimetorum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Parasitus fimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA; Michigan; Canada: Alberta; Iceland, British Isles, Netherlands, Belgium, Germany, Austria, Switzerland, Italy (type locality); Poland; former USSR, including Western Siberia; Iran (beehives); China: Yunnan Prov.; Australia (Gu et al., 1987; Richards 1976; Mosaddegh, 1997; Wallace and Holm, 1985). 7363812ede684713a059042fc0599eaa taxon_parasitus_fimetorum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Parasitus fimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations (only North American records). ; Bombus (Bombus) terricola Kirby, 1837 (as occidentalis); Bombus (Cullumanobombus) rufocinctus Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Fervidobombus) pensylvanicus (De Geer, 1773) (as americanorum) (first record); Bombus (Pyrobombus) bifarius Cresson, 1878; Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Subterraneobombus) appositus Cresson, 1878; (Richards and Richards, 1976) 7c70da974810f4c9aede32ee22e129e2 taxon_parasitus_fimetorum http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Parasitus fimetorum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Parasitus fimetorum is a common inhabitant of compost, rotting vegetation, manure, dung, and subterranean nests of small mammals, and also occasionally bumblebee, bird nests, and beehives. Deutonymphs are phoretic on coprophagous and necrophagous beetles, Aphodius, Geotrupes (Coleoptera: Scarabaidae), Nicrophorus (Silphidae), and Atholus (Histeridae). In the United States, Canada, and Europe, this species was found in bumbleebee nests, but not on the bees. Thus, it likely does not have any specific relationships to bumblebees (Hyatt, 1980; Richards, 1976). b68f31a00ead3e9cd693b45b1e1720b0 taxon_pneumolaelaps http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Pneumolaelaps http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. There are different opinions on the year of the original description of Pneumolaelaps Berlese, e.g., 1920 (Vitzthum, 1930; Casanueva, 1993); 1921 (Hunter, 1966; Hunter and Husband, 1973), or 1925 (Karg, 1984). We checked the original publication and note that it has been published in 1920. The title page of the journal indicates 1921 as the publication year, while the footnote at the end of the article says "Gli estratti di questa Memoria furono pubblicati il 10 Settembre 1920". e9d5af68901fd8b7cbf0e4f792d576ff taxon_pneumolaelaps http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Pneumolaelaps http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription Species of the genus Pneumolaelaps are common inhabitants of nests of bumblebees. They may also occasionally occur in beehives and in subterranean nests of small mammals (Bregetova, 1977; Haitlinger, 1983; Crozier, 1989). The mites may be found in great numbers in active bumblebee colonies, moving rapidly over the nest floor, nectar pots, and brood cells where they feed on nectar and pollen. Female mites become phoretic on bumblebee queens in old nests, and overwinter with the queens in sheltered sites. Feeding behavior is only known for Pneumolaelaps longanalis. This genus is known from the Holarctic, Oriental and Neotropical regions (Eguaras et al. 1997; Evans and Till, 1966; Hunter and Husband, 1973; Karg, 1982, 1984; Rosario, 1981; Royce and Krantz, 1989).
There are, at present, no comprehensive definitions of this genus. Many authors consider it as a subgenus of the genus Hypoaspis (Bregetova, 1977; Evans and Till; 1966; Karg, 1982) while others consider it as a distinct genus (Casanueva, 1993). According to Evans and Till (1966), this taxon has the following apomorphic character states: the stimata are large and peritremes broad, the opistogaster is usually hypetrichous, and there are two ventral setae on genu IV. Karg (1982) defines it based on the following characters: the idiosomal setae filiform, without extreme difference in lengths, unpaired setae frequently present on the posterior part of the dorsal shield; tectum smooth; rows of the hypognathal groove usually with 2 to 6 denticles; the epigynial shield with one pair of setae; 1-2 pairs of setae between the epigynial and anal shields. The latter concept allows the inclusion of free-living species, making the genus, in our opinion, artificial. We accept the point of view of the former authors restricting the genus only to bumblebee-associated species. With the absence of a robust phylogeny of the genusHypoaspis and proofs of the Pneumolaelaps monophyly, this is highly speculative, however.
Qinghailaelaps marmotae, the type species of the genus Qinghailaelaps Gu and Yang, 1984 has been described from Marmota himalayana (Hodgson, 1841) (Rodentia: Sciuridae); another species of this genus, Qinghailaelaps gui Bai, 1992, was found phoretic on a bumblebee (Bombus). Because the genus-level characters of Qinghailaelaps correspond to those of Pneumolaelaps, we consider the former as a junior synonym. The following new combinations are proposed: Pneumolaelaps cavicolous (Gu, Liu et Niu, 1997) from Qinghailaelaps ("cave of bird" in China); Pneumolaelaps gui (Bai, 1992), comb. n. from Qinghailaelaps; Pneumolaelaps marmotae (Gu et Yang, 1984), comb. n. from Qinghailaelaps; Pneumolaelaps qinghaiensis (Li, Yang et Wang, 1998), comb. n. from Qinghailaelaps.

Old World species of Pneumolaelaps associated with bumble bees (Bombus spp.)
Pneumolaelaps bombicolens (Canestrini, 1884) (Poland, former USSR, England, Italy, New Zealand, China)
Pneumolaelaps breviseta (Evans and Till, 1966) (England, New Zealand)
Pneumolaelaps fuscicolens (Oudemans, 1903) (Western Europe, former USSR)
Pneumolaelaps colomboi (Evans & Till 1966) (England, Poland, former USSR)
Pneumolaelaps groenlandicus (Trägårdth, 1904) (Greenland, Russia, Canada)
Pneumolaelaps hyatti (Evans & Till, 1966) (Greenland, England, former USSR, China)
Pneumolaelaps marginepilosus (Sellnick, 1938) (England, Austria, Iceland, Poland, former USSR)
Pneumolaelaps minutissima (Evans and Till, 1966) (England)
Pneumolaelaps qinghaiensis (Li, Yang et Wang, 1998), comb. nov. from Qinghailaelaps (China, ex Bombus) 857d5cb054be7ca7f3b6c1b058b96f4c taxon_pneumolaelaps_aequalipilus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps aequalipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Georgia (type locality); Florida (first record); Illinois (first record); Lousiana (first record); South Carolina (first record); Taxes (first record); Kansas (first record). af4b5ff9c221f9ab3a30fd11c0d16f2c taxon_pneumolaelaps_aequalipilus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps aequalipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Pyrobombus) impatiens Cresson, 1863.; Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (type host); Bombus (Psithyrus) variabilis (Cresson, 1872);
cf9c74acd03a74d98bb30536291bae01 taxon_pneumolaelaps_connieae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps connieae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Michigan (type locality); Georgia. 66705c4634909615f7f491dec6617824 taxon_pneumolaelaps_connieae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps connieae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Bombus) terricola Kirby, 1837 (including occidentalis); Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (as americanorum) (type host); Bombus (Pyrobombus) impatiens Cresson, 1863.; Bombus (Separatobombus) griseocollis (DeGeer, 1773)
547ef0487808dd7159a5a789addfd712 taxon_pneumolaelaps_costai http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps costai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Michigan (type locality); Ohio (first record).
d1076f4993075e62fc68813dfd19f421 taxon_pneumolaelaps_costai http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps costai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Bombus) terricola Kirby, 1837 (including occidentalis); Bombus (Fervidobombus) fervidus (Fabricius, 1798) (first record); Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (as americanorum) (type host); Bombus (Pyrobombus) impatiens Cresson, 1863.
ef4aced7ece68095757a1b5255fb3a9e taxon_pneumolaelaps_groenlandicus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Pneumolaelaps groenlandicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Paralectotypes of Hypoaspis bombicolens var. groenlandica are represented by at least three species of Pneumolaelaps: 1) Pneumolaelaps sp. (single female in bad shape; neither resembles the original drawings nor subsequent interpretation of this species by Bregetova (1977) and Karg (1984)); 2) Pneumolaelaps groenlandica sensu Hunter et Husband, 1979 (we studied two paralectotype females, they do not correspond Trägårdth's original drawings); 3) Pneumolaelaps patae Hunter et Husband, 1979 (1 paratype female from Trägårdth's syntypes; probably, this is species that has been depicted by Trägårdth). Because Hunter et Husband (1979) designated a female of the second species as the lectotype for the Trägårdth's variety, it becomes the unique bearer of the name of Hypoaspis bombicolens var. groenlandica Trägårdth, 1904 and the standard for its application (ICZN, 1999, article 74.1). 2898fbbb512356fc42882835592e4fc6 taxon_pneumolaelaps_groenlandicus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps groenlandicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Greenland (type locality); also recorded for Russia: Jinretlen (East Chukotka) (Hunter and Husband, 1979; Karg, 1984; Trägårdth, 1904). 2ce477590059886dac931a9d50187075 taxon_pneumolaelaps_groenlandicus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps groenlandicus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus sp. (type host)
Bombus (Alpinobombus) hyperboreus Schönherr, 1809
(Hunter and Husband, 1979; Karg, 1984; Trägårdth, 1904).
4d037a2d391e1253ccacbdf53b1bc414 taxon_pneumolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Kansas, Michigan (type locality), Ohio (first record), Pacific Northwest of the United States; Canada: Alberta (Hunter et Husband, 1973; Royce and Krantz, 1989; our data) 0d46adfa91429dc69f486cd2435f1b0b taxon_pneumolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758; Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) affinis Cresson, 1863; Bombus (Bombus) terricola Kirby, 1837 (including occidentalis); Bombus (Fervidobombus) fervidus (Fabricius, 1798) (including californicus); Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (as americanorum); Bombus (Fraternobombus) fraternus (Smith, 1854); Bombus (Psithyrus) suckleyi Green, 1860; Bombus (Pyrobombus) bifarius Cresson, 1878; Bombus (Pyrobombus) bimaculatus Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) impatiens Cresson, 1863 (first record); Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837; Bombus (Pyrobombus) vagans Smith, 1854; Bombus (Separatobombus) griseocollis (DeGeer, 1773) (type host)
41864835bc1121105d8b7b3b30725a40 taxon_pneumolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Pneumolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology The following account on the feeding behavior of Pneumolaelaps longanalis is modified from Royce and Krantz (1989). Wandering P. longanalis congregates in considerable numbers on the brood cells, and larvae feeding on pollen grains provided by the adult bumblebees. When an individual pollen grain was chosen, the gnathosoma was depressed ventrally relative to the opisthosoma, and the grain removed from its sticky substrate by the palps and chelicerae. The nectar-coated grain was held in the region of the hypostome under the encircling palpi and rapidly rotated with the help of the chelicerae. During the manipulation, the nectar coating applied earlier by the foraging bee was stripped from the pollen grain surface. The mite probably also removed much of the surface pollenkitt, a lipoidal substance that covers the pollen grain surface and is incorporated to some extent in the outer wall (exine). Pollenkitt, however, does not trigger feeding in the absence of nectar. It appears that the removed nectar and pollenkitt are dissolved by salivary secretions that are directed over the rotating pollen-grain surface, and then recaptured and redirected to the preoral cavity via the fluid transport system formed by the capitular groove and overlying tritosternum. Following feeding, the stripped pollen grain was discarded and another quickly chosen for manipulation. Females were found to process pollen grains twice as rapidly as either males or deutonymphs. The mites may rupture thin-walled pollen grains during the feeding process, and it is possible that nutrients are acquired from the pollen core in this way. 76765b7fbbed48e67b9e3162f1c8ee76 taxon_pneumolaelaps_longipilus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps longipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Georgia (type locality); Michigan (first record).

fc49032662a4748dc075422dce48e6da taxon_pneumolaelaps_longipilus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps longipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (as americanorum) (type host)
Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus)
Bombus (Psithyrus) variabilis (Cresson, 1872).
d88cd7c578b9675d90c038d61036b42b taxon_pneumolaelaps_mistipilus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps mistipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Georgia (type locality); Michigan.
c355d2c569842bdf45f357392ff50977 taxon_pneumolaelaps_mistipilus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps mistipilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Bombias) nevadensis Cresson, 1874; Bombus (Bombus) affinis Cresson, 1863; Bombus (Fervidobombus) fervidus (Fabricius, 1798); Bombus (Fervidobombus) pensylvanicus (DeGeer, 1773) (as americanorum); Bombus (Fervidobombus) weisi Friese, 1903 (as Psithyrus laboriosus); Bombus (Fraternobombus) fraternus (Smith, 1854); Bombus (Psithyrus) ashtoni (Cresson, 1864); Bombus (Pyrobombus) bimaculatus Cresson, 1863; Bombus (Pyrobombus) impatiens Cresson, 1863 (type host); Bombus (Pyrobombus) perplexus Cresson, 1863; Bombus (Separatobombus) griseocollis (DeGeer, 1773)
50a177577036c8b766a70ec3d989a698 taxon_pneumolaelaps_patae http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Pneumolaelaps patae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Bregetova's (1977) identified mites of the genus Pneumolaelaps from Yakutiya and Chukotka without a striated sternal shield as Hypoaspis bombicolens var. groenlandica, and this interpretation was subsequently accepted by Karg (1982, 1984). Because all of the Trägårdth's syntypes [paralectotypes] we have seen have a distinctly striated sternal shield, we believe that Hypoaspis (Pneumolaelaps) groenlandicus sensu Bregetova, 1977 is, probably, a new species. 275613d981e7edad9d338a190ae50fb6 taxon_pneumolaelaps_patae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps patae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Nunavut; Greenland: Vestgrønland (type locality). 8a41750ab0bd0cfd9151c8060803fbac taxon_pneumolaelaps_patae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps patae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus sp. (type host)
Bombus (Alpinobombus) hyperboreus Schönherr, 1809
Bombus (Alpinobombus) polaris Curtis, 1835
a88693d97b3a2c43e911cdcc353a4a93 taxon_pneumolaelaps_richardsi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps richardsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Alberta (type locality); USA: Michigan.

6f1f3efcfff69bd339048afbd23a9ba7 taxon_pneumolaelaps_richardsi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps richardsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Bombus (Psithyrus) suckleyi Green, 1860; Bombus (Pyrobombus) bifarius Cresson, 1878 (type host); Bombus (Pyrobombus) frigidus Smith, 1854; Bombus (Pyrobombus) huntii Greene, 1860; Bombus (Pyrobombus) mixtus Cresson, 1878; Bombus (Pyrobombus) ternarius Say, 1837
5a8f7ac4829511a84d013204a238caba taxon_pneumolaelaps_sinhai http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pneumolaelaps sinhai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Michigan; Canada: Alberta (type locality); Nova Scotia. 75fccf233b21c41574393c78257c1944 taxon_pneumolaelaps_sinhai http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pneumolaelaps sinhai http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus); Bombus (Psithyrus) insularis (Smith, 1861); Bombus (Psithyrus) suckleyi Green, 1860 (type host); Bombus (Pyrobombus) flavifrons Cresson, 1863; Bombus (Pyrobombus) frigidus Smith, 1854
e3d8780b22d30fc08a13e6d8713d69e5 taxon_podapolipidae http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Family Podapolipidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The genitive case of the Latin word polipus (variant of polypus) is polipi (polypi), thus the family name should be Podapolipidae, not Podapolipodidae. 833cf98955a5f8d0022f75c881f0f1a0 taxon_podapolipidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Podapolipidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes 30 genera and 239 species (Zhang et al., 2011) of internal or external arthropod parasites which often display reductions in numbers of legs and gnathosomal structures. One species, Locustacarus buchneri, is associated with bees.
2973d4347ddc881ac771d7eb6932b2cf taxon_proctolaelaps http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Proctolaelaps http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus Proctolaelaps includes nearly 90 described species. They occur in association with bark beetles, bumblebees and other insects, in nests of birds and mammals, in flowers visited by hummingbirds, in various decaying organic substances, or are synanthropic. Some species feed on small arthropods (including acarid mites), nematodes, fungi, or pollen (Nawar, 1992). Adults of other species are phoretic on insects, although Proctolaelaps nauphoetae (Womersley, 1956) is parasitic in all instars on cockroaches (Egan and Moss, 1969). Many species are cosmopolitan (Halliday et al., 1998).
Five species were recorded in association with nests of bumblebees or phoretic on the adult insects in Europe, Siberia, and Kuril Islands (Davydova, 1988; Karg, 1971; Klimov, 1998; Westerboer, 1963). All of them, except for Proctolaelaps sibiriensis (Davydova, 1988), were found in North America. Proctolaelaps pygmaeus, a widespread species living in a variety of habitats, including honeybee hives, is also given in the key below. In addition, Proctolaelaps bickleyi (Bram, 1956), Proctolaelaps scolyti Evans, 1958, and several unidentified Proctolaelaps species were reported from beehives in Iran (Kamali et al., 2001). da90d7b30d756c316c6b312755a85c31 taxon_proctolaelaps_bombophilus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Proctolaelaps bombophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Our mites well agree with the original description (Westerboer, 1963), except for movable digit of chelicera with one tooth (two in P. bombophilus from Europe) and apices of corniculi less pointed. The species is identified tentatively. Comparison with European material is necessary. d64be1568cb11bf0bf5514128dc49bd6 taxon_proctolaelaps_bombophilus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctolaelaps bombophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: Alberta (first record); USA: Michigan (first record); Germany (type locality); Sweden (first record); Russia: Moskovskaya Oblast', Western Siberia, Kuril Islands (Bregetova, 1977; Davydova and Bogatyrev, 1990; Klimov, 1998; Westerboer, 1963; our data)
867bed1d9ad7fa5f168abb23d02d442f taxon_proctolaelaps_bombophilus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctolaelaps bombophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
North America:
Bombus (Bombus) terricola Kirby, 1837 (as occidentalis)
Bombus (Fervidobombus) fervidus (Fabricius, 1798) (as californicus)
Europe: Bombus sp.
Bombus (Bombus) albocinctus Smith
Bombus (Confusibombus) confusus Schenck, 1859 (as "conficius") (first record). 730de56cf9c9415479f2e7b466805d9d taxon_proctolaelaps_bombophilus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctolaelaps bombophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species is found so far in nests of bumblebees, and also in nests of rodents, including squirrels (Bregetova, 1977; Westerboer, 1963). One specimen was collected phoretic on a bumblebee (our data). Mites from North America were found on brood and dead female inside nest.
9f97fb1adb65729781afe1c5318d7c84 taxon_proctolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA (first record): Michigan; Germany (type locality); Ukraine (Bregetova, 1977; Westerboer, 1963). 27db5841ea88adb9783e874a2d5b0a7a taxon_proctolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
North America:; Bombus (Fervidobombus) pensylvanicus (De Geer, 1773) (as americanorum) (first record); Bombus (Fervidobombus) fervidus (Fabricius, 1798) (first record); Bombus (Pyrobombus) bimaculatus Cresson, 1863 (first record); ; Palaearctic region:; Bumblebee nest, rodent nest (Bregetova, 1977; Westerboer, 1963).
2421f55063ad465a3564ac4ce217b07b taxon_proctolaelaps_longanalis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctolaelaps longanalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species was known only from bumblebee and rodent nests in Europe (Bregetova, 1977; Westerboer, 1963). We have seen a small series of females collected from adult bumblebees in North America. 0968892d561b1ea42c84e3476d3fd3d3 taxon_proctolaelaps_longisetosus http://purl.org/dc/dcmitype/StillImage image/jpeg Proctolaelaps longisetosus, phoretic deutonymph, ex Bombus fervidus, USA: Ohio, BMOC 02-0706-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Proctolaelaps_longisetosus_BMOC_02-0706-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 77d73ebf815863262b5f8baf2296cd00 taxon_proctolaelaps_longisetosus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Proctolaelaps longisetosus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Type host is not indicated in the original description. a01ad077257508a74c02ca6544a26567 taxon_proctolaelaps_longisetosus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctolaelaps longisetosus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution North America:
Apis mellifera Linnaeus, 1758
Bombus (Bombias) auricomus (Robertson, 1903) (first record)
Bombus (Bombus) affinis Cresson, 1863 (first record)
Bombus (Fervidobombus) fervidus (Fabricius, 1798) (first record)
Bombus (Fervidobombus) pensylvanicus (De Geer, 1773) (as americanorum) (first record)
Bombus (Pyrobombus) impatiens Cresson, 1863 (first record)

Palaearctic region:
Bombus (Bombus) lucorum (Linnaeus, 1761)
Bombus (Bombus) terrestris (Linnaeus, 1758)
Bombus (Megabombus) hortorum (Linnaeus, 1761)
Bombus (Megabombus) ruderatus (Fabricius, 1775)
Bombus (Melanobombus) lapidarius (Linnaeus, 1758)
Bombus (Pyrobombus) hypnorum (Linnaeus, 1758)
Bombus (Pyrobombus) pratorum (Linnaeus, 1761)
(El-Banhawy and Nasr, 1986; Karg, 1971; Westerboer, 1963; our data) 64d404f5f20219f664240bc39cccb60f taxon_proctolaelaps_longisetosus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctolaelaps longisetosus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species is found on adult bumblebees and their nests (see below) and in subterranean nests of rodents (Bregetova, 1977; Westerboer, 1963). b478ea6204a296169a65316ab55f652e taxon_proctolaelaps_ornatus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Proctolaelaps ornatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Type host is not indicated in the original description. a5daa9b963715b775c5c189355a24a5b taxon_proctolaelaps_ornatus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctolaelaps ornatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Michigan (first record), Ohio (first record); Canada (Alberta) (first record); Germany (type locality); Ukraine; Russia: Kuril Islands (Bregetova, 1977; Davydova and Bogatyrev, 1990; El-Banhawy and Nasr, 1986; Klimov, 1998; Westerboer, 1963) dd61e24482097f9787e678830599536f taxon_proctolaelaps_ornatus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctolaelaps ornatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
North America:; Bombus (Fervidobombus) fervidus (Fabricius, 1798) (first record); Bombus (Fervidobombus) pensylvanicus (De Geer, 1773) (as americanorum) (first record); Bombus (Pyrobombus) bifarius Cresson, 1878 (first record); Bombus (Pyrobombus) bimaculatus Cresson, 1863 (first record); Bombus (Pyrobombus) ternarius Say, 1837 (first record); Bombus (Pyrobombus) vagans Smith, 1854 (first record); Bombus (Subterraneobombus) borealis Kirby, 1837 (first record); ; Palaearctic region:; Bombus (Bombus) terrestris (Linnaeus, 1758); Bombus (Melanobombus) lapidarius (Linnaeus, 1758); Bombus (Bombus) lucorum (Linnaeus, 1761); Bombus (Megabombus) hortorum (Linnaeus, 1761); Bombus (Pyrobombus) hypnorum (Linnaeus, 1758); Bombus (Megabombus) ruderatus (Fabricius, 1775); Bombus (Bombus) albocinctus Smith; Bombus (Megabombus) tichenkoi (Skorikov, 1925); (Klimov, 1998; Westerboer, 1963; our data);
8e113b2fc8d9f88c3d52c0e4d13293ae taxon_proctolaelaps_ornatus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctolaelaps ornatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species is found on adult bumblebees and their nests (see below) and in subterranean nests of rodents (Bregetova, 1977; Westerboer, 1963). 3ff84673da8fc2075a11f456a92eea60 taxon_proctolaelaps_pygmaeus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctolaelaps pygmaeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA; Canada; localities outside North America include Palaearctic, Oriental, Afrotropic, Neotropic, and Australian regions (Casanueva et al., 1994; Bregetova, 1977; Haragsim et al., 1978; Karg, 1971; Raut and Bhattacharya, 1999; Westerboer, 1963). 477700f613a872e91b42d8909385f531 taxon_proctolaelaps_pygmaeus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctolaelaps pygmaeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Proctolaelaps pygmaeus was collected in a very wide range of habitats, including soil, stored food, bulbs, laboratory insect cultures, plant roots, decomposing plant material, under bark, in subterranean nests of small mammals, birds and in beehives. It may be predatory on small arthropods, but is also a fungivore, and unlike most ascid mites, it is able to ingest solid matter. There is circumstantial evidence that it can attack humans and that the results can be severe enough to cause extensive papular dermatosis (Halliday et al., 1998; Haragsim et al., 1978). In North America, this mite has not been found on bees or in bee nests.
de7da2df6066625748cbe4a9cd0baae2 taxon_proctotydaeus http://purl.org/dc/dcmitype/StillImage image/jpeg Proctotydaeus therapeutikos (Tydeidae): female, dorsal view, ex Scaptotrigona postica, Brazil (modified from Flechtmann and Camargo, 1979) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Proctotydaeus_therapeutikos.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1134b6029a1c1a56a93c4d6f4453f438 taxon_proctotydaeus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Proctotydaeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes 17 species grouped in four subgenera (number of species in parenthesis): Proctotydaeus (4); Neotydeolus Flechtmann et Camargo, 1979 (3); Proctotyduls Kazmierski, 1998 (6); and Oriolella Baker, 1968 (4).
Species of the nominal subgenus are associated with locusts (Acrididae, Orthoptera). The subgenus Oriolella includes species that have been found in barn straw, nests of hawks and owls or on beetles (Coleoptera: Ipidae, Cucujidae). Mites of the subgenus Proctotydulsare found on different plants, straw, ipid beetles (Ipidae), or on noctuid moths (Lepidoptera: Noctuidae). All species of the subgenus Neotydeolus are associated with nests of stingless bees. (Kazmierski, 1998b; Khaustov, 1997). 0a371e4a2a113b40a0888d7697a0746b taxon_proctotydaeus_neotydeolus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Proctotydaeus (Neotydeolus) http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Femur IV not divided. Propodosomal bothridia clavate. Genital papillae absent. Paraproctal suckers (terminal opisthosomal suckers) poorly developed. Tectal setae on tarsus I approximately twice as long as tarsus I; Setae u I minute, spiniform. Males without bifurcate setae, spur on femur IV absent (Kazmierski, 1998b). 8188d704beeeadde6be97d0706e6169a taxon_proctotydaeus_neotydeolus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Proctotydaeus (Neotydeolus) http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology The subgenus includes three species that feed on fungi in nests of stingless bees of the genera Scaptotrigona, Partamona, and Melitoma (Apidae: Meliponini) and, thus, are mutualistic (Flechtmann et Camargo, 1979; Kazmierski, 1998b; Rosa et al., 1985; Rosa, Flechtmann, 1983). 2d3d76c55caa43577644a921a62ca39e taxon_proctotydaeus_neotydeolus_alvearii http://purl.org/dc/dcmitype/StillImage image/jpeg Proctotydaeus (Neotydeolus) alvearii, female, dorsal view, ex Melipona seminigra (modified from Rosa et al., 1985) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Proctotydaeus_alvearii.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 917339d97750924bc048a94b5e75f2d8 taxon_proctotydaeus_neotydeolus_alvearii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctotydaeus (Neotydeolus) alvearii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Maranhão. 7eba66fd6d81692684cc8455b3af3140 taxon_proctotydaeus_neotydeolus_alvearii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctotydaeus (Neotydeolus) alvearii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Melipona seminigra Friese, 1903 (as "Melipona pernigra Moure & Kerr"=Melipona seminigra pernigra Moure & Kerr, 1950) (Apidae: Meliponini). a52fa8b7bb2e5c94d53ed6621d19a30d taxon_proctotydaeus_neotydeolus_alvearii http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctotydaeus (Neotydeolus) alvearii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the nest of Melipona seminigra (Apidae: Meliponini). 7244194a1871d237ccff595c409524ce taxon_proctotydaeus_neotydeolus_partamonae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctotydaeus (Neotydeolus) partamonae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Roraima. 7e0d7925337415ada3ad5ce0f34490ae taxon_proctotydaeus_neotydeolus_partamonae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctotydaeus (Neotydeolus) partamonae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Partamona pearsoni (Schwarz, 1938) 4678ee2ecc7b1e48db1644891fc634b4 taxon_proctotydaeus_neotydeolus_partamonae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctotydaeus (Neotydeolus) partamonae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the nest of Partamona pearsoni (Schwarz, 1938) (Apidae: Meliponini). cf13bfe57fed30000a8c340ae5e20f61 taxon_proctotydaeus_neotydeolus_therapeutikos http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Proctotydaeus (Neotydeolus) therapeutikos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: São Paulo. 450a129a35dcbd4b8a0e5485641c5015 taxon_proctotydaeus_neotydeolus_therapeutikos http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Proctotydaeus (Neotydeolus) therapeutikos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Scaptotrigona postica (Latreille, 1807). 914996b69d45b8dfb55ca2be1dec4593 taxon_proctotydaeus_neotydeolus_therapeutikos http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Proctotydaeus (Neotydeolus) therapeutikos http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Feeds on fungi in the nests of Scaptotrigona postica (Latreille, 1807). Loss of the bee brood decreases from 50% to 3% if the mite is present in the nest (Flechtmann and Camargo, 1979).
49a05d7412477be15360ec97da162c14 taxon_pyemotes http://purl.org/dc/dcmitype/StillImage image/jpeg Pyemotes tritici (Lagrèze-Fossat and Montané, 1851). Photograph by Eric Erbe; digital colorization by Chris Pooley. Identification by Ronald Ochoa. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_tritici.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3c15f8b329c2ad41417d6fef024b109a taxon_pyemotes http://purl.org/dc/dcmitype/StillImage image/jpeg Physogastric females of the mite Pyemotes sp. (Pyemotidae) parasitizing pupa of the bee Hoplitis albifrons argentifrons (Hymenoptera: Megachilidae) from Colorado http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_on_Hoplitis.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 66f6c37a565a4b31c48587d19f258add taxon_pyemotes http://purl.org/dc/dcmitype/StillImage image/jpeg Physogastric female of the mite Pyemotes sp. ex Hoplitis sp. (Hymenoptera: Megachilidae), USA: Colorado http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/PyemotesB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 035cce188279b1d0052fe4311a0d514f taxon_pyemotes http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Pyemotes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Some authors erroneously cite the year for this genus as "1862" and the type species as "Heteropus ventricosus Newport, 1850" (Khaustov & Perkovsky, 2010) e839ec72a41fda436519a5bfdf2a337e taxon_pyemotes http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Pyemotes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus Pyemotes includes polixenous or monoxenous insect parasites exhibiting a wide variety of morphological polymorphisms. Some species of Pyemotes are natural enemies of forest insects or stored product insects. The genus is divided into two groups, scolyti and ventricosus (Cross et al., 1981).
Species of the scolyti group are phoretic on bark beetles (Scolytidae). At least one female morph is a phoretomorph. Phoretomorphic females are shorter than "normal" females and much broader, with thickened legs and enlarged claws. There are also two morphs in the male of Pyetomotes dimorphus Cross et Moser, 1975: "normal" and heteromorphic. The latter morph is distinctly larger, with many greatly enlarged setae, and sometimes setae are duplicated. The mites feed only on bark beetle brood or on immatures of other beetles in the subcortical habitats; none feeds on adults. They are relatively "venomless" and the host remaining alive for about a day after the initial attack (Cross and Moser, 1975).
Species of the ventricosus group are not known to be phoretic, and thus females are monomorphic. At least some species possess venom. The mites inject venom into prey, which causes paralisis and eventual death (Krczal, 1957). The host range includes a variety of hosts, and some species even attack and feed on pupae and adult insects. The bites of Pyemotes tritici (Lagrèze-Fossot et Montané, 1851) can cause severe dermatitis on people handling infested material such as hay. Contact with this mite can also produce asthma or nausea.
Records of Pyemotes from bees include Pyemotes ventricosus (Newport, 1850) from Anthophora retusa (Apidae) in England (Newport, 1850), Pyemotes anobii Krczal, 1957 from a colony of the European honey bee, Apis mellifera (Apidae) in the United States (Cross and Moser, 1975), Pyemotes beckeri Krczal, 1957 (as ventricosus) from laboratory cultures of megachilid bees (Krombein, 1967), and Pyemotes herfsi (Oudemans, 1936) from hives of Apis cerana in India (Dinabandhoo and Dogra, 1982). In the latter case, the mites were considered as pests. In Brazil, Pyemotes tritici (LaGrèze-Fossat & Montagné, 1851) can destroy entire colonies of stingless bees (Tetragonisca angustula, Frieseomelitta varia , Melipona subnitida, Melipona asilvai ) and cause skin irritation in beekeepers (Menezes at al., 2009; Kerr at al., 1996; Nogueira-Neto, 1997).
Pyemotes ventricosus, the type species of the genus, is poorly described and probably has been misidentified by subsequent workers. Oudemans (1937) speculated that the fungal disease of the bee larvae reported by Frison (1923) for Anthophora abrupta in the United States, is in fact physogastric females of Pyemotes ventricosus. Cross and Moser (1975) suggested that this species is only known from the original description and has never been recollected. 47b90cbf68d03702089b2322c6f2307a taxon_pyemotes http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Pyemotes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Cosmopolitan 834ed1a26f16c4126e5b5c306425a9d7 taxon_pyemotes http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Pyemotes http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Parasitizes insects of different orders e8a9dcfffd094854321d870fa8319010 taxon_pyemotes_anobii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes anobii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: California, Georgia, Louisiana; Germany (type locality); Denmark (Krczal, 1957; Cross and Moser, 1975). 864660595beaa22e9143acbcab3bab51 taxon_pyemotes_anobii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pyemotes anobii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Originally described from a culture of Anobium punctatum (De Geer, 1774) (Coleoptera: Anobiidae), also reproduced on pupae and larvae of Sitophilus granarius (Linnaeus, 1758) and Sitophilus oryzae (Linnaeus, 1763) (Curculionidae) in Germany (Krczal, 1957). In the United States it was found on Phloeotribus dentifrons (Blackman, 1921) (Scolytidae), Agrilus lecontei Saunders 1871 (Buprestidae), and Apis mellifera (Hymenoptera: Apidae) (Cross and Moser, 1975).; ; Eickwort (1988) suggested that records of P. venticosus attacking honeybees may refer to Pyemotes anobii, see Notes for Pyemotes venticosus. 7f6e25a1fbeb6f4fbc2b2843fe03251a taxon_pyemotes_anobii http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Pyemotes anobii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found parasitizing larvae of a range of Coleopteran hosts (see below), also found in a colony of the European honeybee Apis mellifera in the United States (California) (Cross and Moser, 1975). 17765aad4b58400245e9e29f3f930aff taxon_pyemotes_beckeri http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes beckeri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, Florida, Louisiana, Maryland, Mississippi, New York, and Virginia; Germany (type locality) (Cross and Moser, 1975; Krczal, 1957; Krombein, 1967). 986b6b2dce25f74b2627434457d29f6e taxon_pyemotes_beckeri http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Pyemotes beckeri http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology In the United States this species was recorded from Lyctus planicollis LeConte, 1858 (Coleoptera: Lyctidae) in Mississippi, Scolytus multistriatus (Marsham, 1802) (Scolytidae) in Louisiana, and a wasp nest in Virginia (Arlington) (Cross and Moser, 1975). Krombein (1967) reported "Pyemotes ventricosus" from an array of sphecid and vespid wasps, as well as from nests of megachilid bees infested in the laboratory. Because his material included numerous samples from nests of wasps from Arlington (Virginia), and Cross and Moser (1975) reported Pyemotes beckeri from a wasp nest from exactly the same locality, we assume that Krombein's material in fact was misidentified. Unfortunately, without Krombein's material we can not confirm this assumption or check whether it is represented by one or more species. Unverified records of Pyemotes beckeri from North American Hymenoptera include Tripargilum clavatum (Say, 1837) (Maryland, New York, Virginia), Tripargilum tridentatum tridentatum (Packard, 1867), Tripargilum tridentatum archboldi (Krombein, 1959) (Florida), Trypoxylon frigidum Smith, 1856 (Virginia), Psenulus pallipes parenosas (Pate, 1944) (Virginia) (Sphecidae), Stenodynerus krombeini Bohart, 1953 (New York), Euodynerus foraminatus apopkensis (Robertson, 1901) (Virginia) (Vespidae), Prochelostoma philadelphi (Robertson, 1891) (Virginia, infested in laboratory), Aschmeadiella bigeloviae (Cockerell, 1897) (Arizona, infested in laboratory), Aschmeadiella bucconis denticulata (1878) (Arizona, infested in laboratory), Osmia pumila Cresson, 1864 (infested in laboratory, the nest is probably from Maryland) (Megachilidae) (Krombein, 1967).
Outside North America, this species was found on larvae of Anobium punctatum (De Geer, 1774) (Coleoptera: Anobiidae), Sitophilus granarius (Linnaeus, 1758), and Sitophilus oryzae (Linnaeus 1763) (Curculionidae) in Germany (Krczal, 1957).
Krombein (1967) noted that "normal" females move relatively rapidly. They are able to leave an infested nest after the host insects have been killed, and enter adjacent nests through the breached entrance plugs and cell partitions. The mites attack eggs, larvae, pupae, as well as paralyzed prey stored by wasps (insects and spiders). This species of Pyemotes can cause serious problems in laboratory settings, as mite females can easily infest cultures of bees through the split halves of the trap nests. 89d7b1419f32d4b5767c88557a4b4e6c taxon_pyemotes_herfsi http://purl.org/dc/dcmitype/StillImage image/jpeg Pyemotes herfsi, female, ventral and dorsal view, ex Tineola bisselliella (after Oudemans, 1936). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_herfsi_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e279a7e29d01e0addedf4307187f804b taxon_pyemotes_herfsi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes herfsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Holarctic and Oriental regions (Oudemans, 1936; Dinabandhoo and Dogra 1980; Broce, 2006; Marutani et al., 1992). (type locality - Gemany: Leverkusen) a7828e3230d3f689cf813a49686f98be taxon_pyemotes_herfsi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pyemotes herfsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Many insect species from different orders, including Apis cerana Fabricius, 1793. Type host - Tineola bisselliella (Hummel, 1823) (as Sitotroga biselliella Hum.) 3130be14739b457b8e0a3dc4cceb5079 taxon_pyemotes_herfsi http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Pyemotes herfsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This mite parasitizes a variety of insect hosts, including bees. Dinabandhoo and Dogra (1980) report that colonies of Apis cerana infested by Pyemotes herfsi in Kulu, India show symptoms of restlessness or lethargy and do not produce brood. The infested colonies were recovered by fumigating them with dichlorvos. Pyemotes herfsi causes high incidences of red, itching, and painful welts on people in the midwestern United States, where the mite is normally preying on gall-making midge larvae on oak leaves (Broce, 2006).
The below account on biology is based on Herfs (1926) and Cross (1965).
Both males and females pass through all immature stages while still within the enormously distended, globular opisthosoma of the mother mite, and are born as adults. Most males are born during the first half of the birth period and normally are present only in small numbers (three to five percent of the total number of offspring at 25°C). They seldom leave the body of the mother and are parasitic upon it. At 25°C, males of P. herfsi removed from the maternal "opisthosomal globe" died 16 to 25 hours after removal, presumably from starvation, and if placed on the bodies of normal, healthy hosts did not feed. Males are usually found resting quietly on the opisthosoma of the mother, for the most part near the vulva. They may assist in the birth of the females by grasping the latter with the hind legs and pulling, shifting the grasp, pulling again, etc. This aid is not necessary since birth of the females continues smoothly in the absence of males although taking slightly longer (average birth time of females of P. herfsi with males, 171 seconds, average time without males, 217 seconds). As soon the newly born female completely leaves the birth canal, the male initiates copulation. In he grasps the female and adjusts, with the assistance of the hind legs, her body until contact is made. This entire process of adjustment and mating may take 10 minutes or more, although actual copulation probably lasts only 20 to 30 seconds.
Males of P. herfsi normally assist only in the birth of the females, paying no attention to emerging males. They likewise ignore mated females, although one or more males may attempt to disrupt pairsin copulo. Early in the birth period of the mother, only a single P. herfsi male is usually found on the opisthosoma but as this period progresses, several may be present. Shortly (within 10 minutes) after being mated, the young female leaves the body of the mother and seeks food. If, however, males are lacking on the maternal hysterosoma, young females do not leave but remain resting upon the latter for some time awaiting the birth of a male. Only after a full day or longer do they leave.
Unmated females seemingly suffer a high mortality, only about 10 percent of those in experimental cultures surviving to produce offspring. Such females are arrhenotokous, producing only males. The total number of offspring produced compares favorably with that of mated females of the same size. According to both Krczal and Herfs, young females die of starvation within 48 hours of their birth if food is not found. This statement makes dispersion and perhaps overwintering difficult to explain since it is obviously the young females which must disperse. Herfs found that young females unable to find a host would sometimes return to the mother and feed in the manner of the male on the parent's hysterosoma. Rarely this filial parasitism became permanent and ended with the death of the mother and the production of offspring by the daughter. More commonly it was quite transitory and resulted in no visible harm to the mother while seemingly providing the daughter with enough food to prevent at least immediate starvation.
Young females of P. herfsi are stimulated by moving objects soon after leaving the mother, stretching the body toward the source of the stimulus. Such behavior indicates dispersion by phoresy, a phenomenon which has been observed only rarely in Pyemotes despite its widespread occurrence in other members of the family. In view of the fact that young females starve so quickly, the questions of their survival between generations of the host and of overwintering remain essentially unanswered. As mentioned by Krczal, perhaps the phoretic period is ephemeral, serving only to bring the females into fortuitous contact with one of a number of possible hosts. This could explain their rare and sporadic occurrence in the nests of solitary bees and social wasps, hosts in which an overwintering period would be required of the young females. Presumably, such a period would result in their migration or death, thus preserving the host from further attack until the next "accidental" infestation occurred.
Upon finding a host, the young female selects a favorable location and inserts her chelicerae, an action which may elicit striking or curling movements from the host. Within a short time, the host begins to show signs of paralysis, this increasing until the larva is completely incapable of movement. Time elapsed between onset of attack and complete paralysis depends upon the number of punctures made, but generally is two to four hours. The same larva may be attacked by several mites, Herfs having counted a maximum of 178 on a single host. The parasite develops quickly, the opisthosoma increasing in both length and width until the mite assumes a form quite similar to an incandescent light bulb. This stage is usually reached in about 24 hours. Complete development, in which the opisthosoma becomes a huge, turgid sphere may be completed in another 24 hours, although it usually requires seven to eleven days to reach maximum size. At 25° C the average time from initial insertion of the chelicerae to birth of the first offspring is 9.5 days in P. herfsi. The size of the opisthosomal globe is well correlated with the number of offspring produced. In cases of multiple parasitism the size of the opisthosoma will be reduced, and females concerned will produce a relatively small number of offspring, sometimes averaging as low as six or seven. Females ofP. herfsi reared singly produced a maximum of 284 offspring and averaged 128 at 25°C and 150 at room temperature.
The birth period, usually terminated by the death of the female, ranged from 9 to 33 days and averaged 17.3 days. The average life span of the females in Herfs' cultures at 25° C. was 26.8 days as measured from the beginning of feeding to the death of the mite. The average life span of the male is much shorter, since it dies with the mother mite. It was found that longevity of P. herfsi males was also positively correlated with amount of sexual activity. The temperature of the microhabitat was also an important factor affecting both longevity and fecundity of both sexes. Several previous workers have found 25 to 30° C to be optimum and Herfs noted a loss in the ability of the mother to give birth at 35°C. 8713c2d58788c77fa1e4b990cf236fa3 taxon_pyemotes_herfsi http://purl.org/dc/dcmitype/Text text/html Management of the Bee Mite Species Pyemotes herfsi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Management Colonies of Apis cerana infested by Pyemotes herfsi in India were cured by three treatments of fumigation with dichlorvos (Dinabandhoo & Dogra, 1980).10 ml of dfchlorvos solution (50 ppm) was applied to a strip of blotting paper stapled to an empty frame. To avoid direct contact by bees, the strip was enveloped by perforated paper. A treatment frame was put into the brood chamber in the spring (3 treatments each of 12 h, at intervals of 2 days) or in the autumn (5 treatments each for 24 h on alternate days). Post-treatment examination revealed that the mite was controlledfollowing three fumigations. No mites were found on bees examined after treatment. The uptake of dichlorvos by the bees was calculated to be 25G-500 ng per colony (5,000-10,000 bees), whtch is considered a safe level (after Dinabandhoo & Dogra, 1979).

5eb4c527d8f9bc83483ddbc25dfc21e4 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Physogastric female of the mite Pyemotes sp. ex Hoplitis sp. (Hymenoptera: Megachilidae), USA: Colorado title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/PyemotesB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9752b760091fa42284df5898128d24b8 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Physogastric females of Pyemotes sp. on pupa of Hoplitis sp. (Hymenoptera: Megachilidae),  USA: Colorado http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_on_HoplitisB.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ba1c18226717752f74aa0f0efac8e00e taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Males are similar to Pyemotes herfsi (Oudemans, 1936) as diagnosed by Cross et al. (1981), p. 182, couplet 14. Our species differs by the fourth propodosomal pair of setae being distinctly thinner than setae d and e (stout, as thick as setae d and e in P. herfsi). 4ee8029471d50dbb85b155328a8e6199 taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Colorado. 8ef70d2c62081a1bbb86b12fe025d935 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Hoplitis fulgida. da16d81c90227d8530ea5f12bb530340 taxon_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found parasitizing pupae of Hoplitis sp. cee501c61d84270c324f9410e10b0930 taxon_pyemotes_sp2. http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes sp2. http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution India: Himachal Pradesh 1b6587a0af198dec99e1b2711152458b taxon_pyemotes_sp2. http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pyemotes sp2. http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Apis mellifera Linnaeus, 1758; Apis cerana Fabricius, 1793; Megachile (Pseudomegachile) flavipes Spinola, 1838 8a38b852a48da0f7aefb53cc22e3c009 taxon_pyemotes_tritici http://purl.org/dc/dcmitype/StillImage image/jpeg Pyemotes tritici (Lagrèze-Fossat and Montané, 1851). Photograph by Eric Erbe; digital colorization by Chris Pooley. Identification by Ronald Ochoa title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_tritici.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 92383ed7a1a05eac78ce574aa0f3eb35 taxon_pyemotes_tritici http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes tritici http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Cosmopolitan (Moser et al. 1987), including USA: Hawaii, Georgia, Louisiana, Kansas; Mexico: Sonora; Cuba (Cross & Moser, 1975); Brazil (Menezes at al., 2009); France (type locality), Iran (Kamali et al. 2001). 4e9ec5a71b7120701af6d3ded2d8a60f taxon_pyemotes_tritici http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pyemotes tritici http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Insects of various orders. In Brazil, the following managed stingless bee species were attacked (Menezes at al., 2009):; ; Tetragonisca angustula (Latreille, 1811); Frieseomelitta varia (Lepeletier, 1836); Melipona subnitida Ducke, 1911; Melipona asilvai Moure, 1971.; ; Eickwort (1988) suggested that records of P. venticosus attacking honeybees may refer to Pyemotes tritici, see Notes for Pyemotes venticosus. b47e7934218454477ee2656b60319ad3 taxon_pyemotes_tritici http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Pyemotes tritici http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology P. tritici, may produce severe skin lesions, asthma, or nausea in humans. It has also been shown to have great potential as a biological control agent for insect pests (Kethley, 1982). Its effectiveness as a parasitoid results from several characteristics (Bruce, 1989):

(1) high reproductive potential;
(2) short life-cycle (4-7 days);
(3) no intermediate hosts or food sources required-- all development occurs within the opisthosomal sac of the adult female and all offspring are born live as sexually mature adults;
(4) only females are parasitic and represent about 90-95% of the population;
(5) females mate immediately at birth and begin host-seeking activity;
(6) populations are easily reared and synchronized in the laboratory; and (7) they are cosmopolitan in distribution.
a8f259e8f61cc47918a1e688ce063440 taxon_pyemotes_tritici http://purl.org/dc/dcmitype/Text text/html Management of the Bee Mite Species Pyemotes tritici http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Management Pyemotes tritici, was reported to kill four colonies of stingless bees Tetragonisca an­gustula and one colony of Frieseomelitta varia in Brazil. The infestation was terminated by isolating the dead colonies from the meliponary (Menezes at al., 2009). ccb11ca6a2f4f811fd15c0e4aaf9d911 taxon_pyemotes_ventricosus http://purl.org/dc/dcmitype/StillImage image/jpeg Pyemotes ventricosus, from nest of Anthophora retusa, England (after Newport, 1853) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_ventricosus.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 634cfd7d59b37bb128f2e1bf456126b4 taxon_pyemotes_ventricosus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Pyemotes ventricosus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution England (see Notes). de38056a477a7e94ea0591779f4c6582 taxon_pyemotes_ventricosus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Pyemotes ventricosus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Anthophora retusa (Linnaeus, 1758) (see Notes) efb30ff4bf1fe100f4a7c3ca5c47445e taxon_pyemotidae http://purl.org/dc/dcmitype/StillImage image/jpeg Pyemotes tritici (Lagrèze-Fossat and Montané, 1851). Photograph by Eric Erbe; digital colorization by Chris Pooley. Identification by Ronald Ochoa.  title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Pyemotes_tritici.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bb0587cc57ccd8142b9c68a5ce5037e7 taxon_pyemotidae http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Family Pyemotidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The family-group name Pediculoididae was proposed by Berlese (1897) for Pediculoides Targioni-Tozzetti, 1878. When it was realized that this genus is a junior synonym of Pyemotes Amerling, 1861, a new family name, Pyemotidae, was proposed (Oudemans, 1937). Although ICZN treats family- and genus-group names independently (i.e., synonymy of a type genus does not affect the validity of the corresponding family-group name), article 40.2 stipulates that substitute family-group names proposed before 1961 is to be maintained if it is in prevailing usage (ICZN, 1999). This is the case for Pyemotidae. The appropriate citation for this family name is Pyemotidae Oudemans, 1937 (1897), see recommendation 40A (ICZN, 1999). a4489aecd15bbe9d24ef4bbd82da94cc taxon_pyemotidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Pyemotidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes three genera according to Lindquist (1986), two genera with 25 species according to Kethley (1982), and one genus and 24 species according to Zhang et al., 2011. One described species is known from the late Eocene (Khaustov and Perkovsky 2010). Adult females of Pyemotidae are parasites of all developmental stages of holometabolous insects, especially subcortical beetles and stored product insects. When the female sucks the haemolymph of the host her posterior opisthosoma becomes greatly swollen because of developing eggs (physogastric female). Eggs hatch within the female, and adults are the only developmental stage in the family. Adult males emerge first, pierce the swollen opisthosoma with their mouthparts and remain near the opening. Males copulate with females as the latter emerge from the mother's body. The copulation triggers searching behavior in females, which immediately leave to attack a suitable host.
The genus Pyemotes Amerling, was recorded parasitizing bees' larvae, pupae, and, raraely, adults. 0b13b76b6f85e73be6c2d974337b061c taxon_roubikia http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Roubikia latebrosa at the entrance of the metasomal acarinarium of bee Tetrapedia sp. (Peru, BMOC-03-1008-052)  http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_latebrosa_BMOC-03-1008-052_acarinarium.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2c4075bf955ca241b0c769b67acf5847 taxon_roubikia http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Roubikia imberba, holotype, Argentina (BMOC 04-0508-252)  http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_imberba_BMOC_04-0508-252.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 5e0d5a774b3bc293aafac6be3388104a taxon_roubikia http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Roubikia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description (unique characters only). Phoretic deutonymph. Gnathosomal solenidion and gnathosomal setae present and free palpi absent. Coxal fields IV closed. Apodemes of ps1 partially fused anteriorly. Dorsal cuticular folds of ambulacra I-III weakly developed, with distal part smaller than proximal.
Larva. Claparède's organ shaft slightly asymmetrical, distinctly narrowing terminally and ending in button-shaped dome.
Adults. Paraxial and antiaxial rutellar lobes (rlp and rpa) fused ventrally, paraxial lobe distinct only dorsally. Supracoxal seta spiniform, with rounded tip. Dorsal condylar plate of femur-tibia joint narrow, posterior.
Female. Spermatophores present. Inseminatory canal cylindrical, well sclerotized, protruding inside spermatheca. Condylophores with short sclerotized portion and distinct proximal unsclerotized portion connected to the tarsus.
Male. Genital setae represented by transparent disk. Genital setae distinctly (more than their diameter at base) anterior to progenital folds. Tarsal setae e III-IV absent. Setae s and w IV separated, w submedial, s subapical. Sclerotized portions of condylophores fused and incorporated into disto-ventral sclerotized tarsal wall, pretarsal suckers not developed. Heteromorphic males present. 8a4622e7554ddbb5b30b79d01d54fb19 taxon_roubikia http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Roubikia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Neotropical region. ebd51babc144755c3a4db97d8a5bfbae taxon_roubikia http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Roubikia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Species of Roubikia are mutualists or commensals feeding on the nest materials, fatty acids from floral oils, or probably, on fungi harmful to the bee larvae (Cordeiro at al., 2010). The presence of a metasomal acarinarium harboring Roubikia latebrosa from Peru (Fig. 1) further suggests mutualistic relationships (Klimov et al., 2007b).
The four described species are exclusively associated with Tetrapedia (Apidae: Tetrapediini). Roubikia panamensis and R. imberba occur on cleptoparasitic bees of the genus Coelioxoides (Apidae: Tetrapediini) attacking their principal host (Alvez-Dos-Santos et al., 2002). Biology is only known for Roubikia panamensis. Inert heteromorphic deutonymph is unknown. fef59112c9bb610c00c2a35bdd88e864 taxon_roubikia_imberba http://purl.org/dc/dcmitype/StillImage image/jpeg Roubikia imberba, phoretic deutonymph, holotype, ex Tetrapedia sp., Argentina (BMOC-04-0508-252), BMOC 04-0508-252 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_imberba_BMOC_04-0508-252.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 845e224526cd7e54f6452d11f1a69a1c taxon_roubikia_imberba http://purl.org/dc/dcmitype/StillImage image/jpeg Tetrapedia sp. with phoretic deutonymphs of Roubikia imberba, Argentina (BMOC-04-0508-252) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_imberba_BMOC-04-0508-252a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3d7063c19928bc9ca52484334c20a2e4 taxon_roubikia_imberba http://purl.org/dc/dcmitype/StillImage image/jpeg Tetrapedia sp. with phoretic deutonymphs of Roubikia imberba, Argentina (BMOC-04-0508-252) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_imberba_BMOC-04-0508-252b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 4bcf8d483d4585a312e309f484f1bd71 taxon_roubikia_imberba http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Roubikia imberba http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina: Tucumán (type locality), Salta. 5bc8bc2bd66099ce49861796c38e23f7 taxon_roubikia_imberba http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Roubikia imberba http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Tetrapedia sp. (type host); also phoretic on Coelioxoides waltheriae Ducke, 1908 and Coelioxoides exulans (Holmberg, 1887), both known cleptoparasites of Tetrapedia species (Apidae: Tetrapediini) (Alves-dos-Santos et al. 2002). 836836ec12380279dda5d7e979eca43b taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Roubikia latebrosa, phoretic deutonymph, holotype, ex Tetrapedia sp., Peru, BMOC 96-0510-213 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_latebrosa_BMOC_96-0510-213.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bcebb2e4c755132d3f70e1f1ed24e223 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Tetrapedia sp. with phoretic deutonymphs of Roubikia latebrosa, Peru (BMOC-03-1008-052) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_latebrosa_BMOC_03-1008-052.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b431337dfd9870807b03d27275ce11aa taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg phoretic deutonymphs of Roubikia imberba near the entrance of the metasomal acarinarium of Tetrapedia sp. (Peru) (BMOC-03-1008-052) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_latebrosa_BMOC-03-1008-052_acarinarium.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1713211819d78db7d6a9ca6094169c4e taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Deutonymph. Gnathosomal setae present, well-developed. Dorsal extensions of apodemes I-II not completely surrounding setae scx. Setae si about twice as long as se or longer, usually as long or longer than distance from lateral edge of prodorsal shield to base of si + distance between si. Genual setae mG I slightly pectinate. Setae mG II nearly as long as leg II (with claw). Other characters as in Roubikia panamensis (Baker, Roubik & Delfinado-Baker 1987).
Notes. Phoretic in the acarinarium situated on the 1st metasomal tergite. 991654066d6b909784d6a3a571ccde24 taxon_ http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription This species can be distinguished from other species of Roubikia by the relatively long internal scapular setae si that are twice as long as se or longer (si less than twice as long as se in all other species of Roubikia known to us). bb24c2f6cbc2b8392b3cb5034ed8a389 taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Peru. 168bb28812383d7407ce518c16fc7b92 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Tetrapedia sp. bdd6dad6cc93c7fe97f8362dd51378a1 taxon_roubikia_officiosa http://purl.org/dc/dcmitype/StillImage image/jpeg Roubikia officiosa, phoretic deutonymph, holotype, ventral view, ex Tetrapedia maura, Mexico, BMOC 97-0331-028 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_officiosa_ventr_BMOC-97-0331-028.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 79c38333eecadd2d48f7bce2c579354d taxon_roubikia_officiosa http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Roubikia officiosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Differs from all known species by the following characters: setae mG I smooth, setae d1 nearly as long as c1, and setae mG II distinctly shorter than combined length of femur-tibia II.
Other instars unknown. dd42c6385f926503d21bca7e40ea2d36 taxon_roubikia_officiosa http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Roubikia officiosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Jalisco 393ccace24e72d8b34f108848c0b2849 taxon_roubikia_officiosa http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Roubikia officiosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Tetrapedia maura. 97e52082c383f030c8a53ed7d27dae1a taxon_roubikia_panamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Roubikia panamensis, phoretic deutonymph, ex Tetrapedia sp., Brazil, BMOC 87-0606-003 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_panamensis_BMOC_87-0606-003.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bf6d32f99c9194fcc43077351bdffaee taxon_roubikia_panamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Coelioxoides waltheriae with phoretic deutonymphs of mite Roubikia panamensis, Bolivia (BMOC 04-0508-245 ) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Roubikia_panamensis_BMOC_04-0508-245.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 25c8340a9d922fa260e7fb79ad3167c5 taxon_roubikia_panamensis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Roubikia panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Probably a complex of species. Specimens from Panama have setae mG II as long as leg II (including claw), while in specimens from Mexico and Brazil, these setae are distinctly longer. 551024eb9f713a8268f64a2eed8eab0a taxon_roubikia_panamensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Roubikia panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Panama (type locality), Mexico: Chiapas, Nayarit, Tamaulipas; French Guiana, Brazil, Bolivia. f620992b18a20ccc48e3a376da573e03 taxon_roubikia_panamensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Roubikia panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations ; Tetrapedia sp. (type host); Tetrapedia diversipes Klug, 1810; Tetrapedia peckoltii Friese, 1899; Coelioxoides waltheriae Ducke, 1908 (cleptoparasite of Tetrapedia diversipes). 7761f5d3b661cd66dbb41923b46240a6 taxon_roubikia_panamensis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Roubikia panamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology In Brazil, mites associated with Tetrapedia diversipes were shown to be beneficial to the bee as the bee mortality rate in nests was inversely correlated to the level of mite infestation (Cordeiro et al., 2010). The mites presumably feed on fungi harmful to the bee larvae inside the bee nests (Cordeiro et al., 2010).
In Panama, Tetrapedia sp. nests primarily from the late dry season until the early wet season (April to July), and occasionally in the early dry season (November to February). The Tetrapedia female collects floral oils to combine with pollen provisions, and gathers dry soil to make partitions between the cells. Both materials are carried on the hairs of the tibial scopae. Soil in the cell partitions appeared to be admixed with a resinous substance and much of it formed small, shiny pellets less than a half of millimeter in diameter. Mature larvae produced fecal pellets, which are about 1.2 mm long and another, cigar-shaped. Several hundred mites were scattered among the two types of pellets. This was the approximate mite abundance in each of three completed nests having 5-6 bee cells. A nest containing young larvae had several mites in the loose soil fill between each cell and also on the pollen provision. The mites presumably fed on materials in the cells, and possibly on the fatty acids from floral oils mixed with some of the fill dirt. Mites were much more apparent in nests from which bees had emerged than in the nest containing young larvae. Adults of Tetrapedia routinely harbor mites on the basal metasomal tergites and hind legs, particularly on the scopae. A characteristic of Tetrapedia is their repeated visitation to small patches of dry soil in which females collect soil for nest construction. The mites apparently disperse to new nests by leaving a nest with emerging bees. An individual mite was seen wandering within a small (10 cm diameter) area in which three female Tetrapedia had been seen collecting loose dirt. The mites thus possibly disperse among female bees at such a restricted foraging spot, and they probably arrive at the nests of more than one apid bee in this manner (Roubik, 1987).
The shape and internal structure of the "nematodes" reported in the female spermathecae (OConnor, 1993a) are consistent with those of astigmatid mite spermatophores (Griffiths & Boczek, 1977) 303de026fa41f760bb4a5cffde1825ee taxon_sancassania_ctenocolletacarus http://purl.org/dc/dcmitype/StillImage image/jpeg Ctenocollectacarus longirostris, phoretic deutonymph, ex Ctenocolletes centralis, Australia, BMOC 86-0406-003 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Ctenocolletacarus_longirostris_BMOC_86-0406-003_dn.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9b68a50399f760f8374a21b2c22b691e taxon_sancassania_ctenocolletacarus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Sancassania (Ctenocolletacarus) http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia. 069ee9e20a580063b10a6ec7234808c0 taxon_sancassania_ctenocolletacarus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Sancassania (Ctenocolletacarus) http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Endemic Australian bees of the genus Ctenocolletes (Stenotritidae). 5e620380eddb82015e9fd7bca06b9971 taxon_sancassania_ctenocolletacarus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Sancassania (Ctenocolletacarus) http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology All stages of mites complete breeding cycle occurs in the hosts' sealed brood cells, except for the phoretic deutonymphs, which can leave the cells and disperse on their bee hosts. The deutonymphs are usually phoretic in specialized circular metasomal pouches on the lateral margins of metasomal terga 3 and 4 (acarinaria). Phoretic deutonymphs are found on 70-90% of four species of female bees. The mites and their hosts are univoltine. One or a few deutonymphs dismount from nesting females into each new blood cell and rapidly transform through non-mobile tritonymphs to males and females. Female mites oviposit on the ceiling of the cell during the host larva's early instars, producing hundreds of larvae and then protonymphs by the time the host larva has completed feeding. Large numbers of mites (nearly 300) develop in each host cell, with no obvious deleterious effects on the developing bee. Larvae and protonymphs aggregate on the host bee larva, possibly to avoid being ingested, and perhaps feeding only on pollen adhering to the lateral surfaces and mouthparts of the bee larva, probably this feeding activity helps the developing bee to keep its integument clean and pathogen free. The mites also convert the pasty feces to a firmer, drier state, which makes it less prone to fungal growth. Non-deutonymphal stages feed upon pollen, host larval feces, and an amorphous substance that may originate from the bee larva's surface or from its feces (after Houston, 1987). 4d0208ea5c28259126dbdf79ae55b318 taxon_sancassania_ctenocolletacarus_brevirostris http://purl.org/dc/dcmitype/StillImage image/jpeg Sancassania (Ctenocolletacarus) brevirostris, phoretic deutonymph, ex Ctenocolletes nigricans, West Australia, BMOC 86-0406-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Ctenocolletacarus_brevirostris_BMOC_86-0406-001_dn.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9971faba4de368c013bfcfec55290e7b taxon_sancassania_ctenocolletacarus_brevirostris http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sancassania (Ctenocolletacarus) brevirostris http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia: Western Australia (incl. Merredin - type locality). e2e87e11e0568855f230cb0e1b98df51 taxon_sancassania_ctenocolletacarus_brevirostris http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sancassania (Ctenocolletacarus) brevirostris http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ctenocolletes albomarginatus Michener, 1965 (type host).; Ctenocolletes nigricans Houston, 1985 79384eccda1e39c00c4fd4bf47e93e77 taxon_sancassania_ctenocolletacarus_brevirostris http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sancassania (Ctenocolletacarus) brevirostris http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Deutonymphs are phoretic in circular metasomal acarinaria on the lateral margins of metasomal terga 3 and 4. See Houston (1987) for a detailed biological account of this species. f7be7b8752218c05827f7eb8b0cf2161 taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Sibling species of C. longirostris. b5ded3bfa14362e1cfd68d08652de88a taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia: Western Australia (type locality). b5e74cf5aee6d9654cd4766806d50b37 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ctenocolletes centralis Houston, 1983 (type host).; Ctenocolletes nicholsoni (Cockerell, 1929) 8bb5fb19aa37e8377e92a1accda4339d taxon_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Deutonymph are found phoretic in circular metasomal acarinaria on the lateral margins of metasomal tergites 3 and 4. Houston (1987) gives a detailed biological account of "C. longirostris" (which may partially include C. grandior). 9cdc2174257e2e5f7212c1a95963721a taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Sibling species of C. grandior. 52418bc0558c7279053f43682f7cc31c taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ctenocolletes nicholsoni (Cockerell, 1929) (type host); Ctenocolletes centralis Houston, 1983 e732349e82aa0f88e0554c8b0f41ec25 taxon_sennertia http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of Sennertia americana, ex Xylocopa virginica, Virginia (BMOC 90-1212-025). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_americana_BMOC_90_1212_025.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 44f4ccc76e923732ead67767561a9f5d taxon_sennertia http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Sennertia segnis on the carpenter bee Xylocopa californica, Arizona (BMOC 04-1222-155). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_segnis_on_host_BMOC_04-1222-155.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e8853e02d2d26ccad022c0c9ba992c6f taxon_sennertia http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Sennertia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description (unique character states only). Phoretic deutonymph. Gnathosomal solenidion present and setae on free palpi absent and free palpi absent. Prodorsal shield absent. Cupules im situated at level of bases of legs III.
Immobile deutonymph absent.
Adults. Cupules im dorsal (correlated with deutonymph).
Male. Main part of progenital sclerites lateral to genital capsule. Progenital sclerites not touching each other. Body of dorsal supporting sclerite, posterior to base of aedeagus absent.
Larva. Claparède's organs absent. 828a33271f451ab038241950c12a2a4e taxon_sennertia http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Sennertia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Worldwide, except for Antarctica. In contrast to Chaetodactylus, there is a clear division between New and Old World lineages. 05f7f05c2010bb15ada61f68204ad9bf taxon_sennertia http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Genus Sennertia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Species of this genus are associated with xylocopine bees Ceratina and Xylocopa (Apidae). Sometimes monophyletic groups of mites occur on monophyletic groups of hosts, indicating their close biological relationships and possible codivergence. Host associations of major groups of Sennertia are recorded here.
The majority of Sennertia disperse as heteromorphic deutonymphs on adult bees, however the Sennertia vaga group does not form deutonymphs and disperses as feeding instars. Reproduction and feeding also probably occur during dispersal.
The interactions of Sennertia with their hosts remain largely unknown. There are conflicting accounts suggesting either negative or neutral effect of the mite presence. In the former case, the damage to developing bees was marginal and always substantially lesser than that of Chaetodactylus. Some species (Sennertia sayutara, Sennertia devincta) are phoretic inside special pouches (acarinaria) on the body of certain Ceratina, suggesting mutualistic relationships. Unfortunately, the biology of such species has not been studied. Several species, such as, Sennertia koptorthosomae, have special pouches filled with fungal spores (sporothecae) on the hysterosoma (Klimov & OConnor, 2008), suggesting that they may transfer fungi from one bee nest to another. 3a9554e65877ed1b829f189f1addf1c1 taxon_sennertia_aldeodadi http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia aldeodadi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Assigned originally to the cerambycina-group cf5ed29ac211d76f1d4f798405b055e8 taxon_sennertia_aldeodadi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia aldeodadi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mauritius: Mahébourg (as Mohebourg) (Haitlinger, 2000) 2bbf29712ef281d1d0bd48cf3b96f719 taxon_sennertia_aldeodadi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia aldeodadi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "undetermined Anthophoridae" (Haitlinger, 2000) afe9929f464a8999e17879741984d866 taxon_sennertia_alfkeni http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia alfkeni, heteromorphic deutonymph, China, ex Xylocopa appendiculata (BMOC 90-1212-001). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_alfkeni_dn_BMOC_90-1212-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8ec6d8572b033ed3f78b88ca7d7edfc0 taxon_sennertia_alfkeni http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia alfkeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan, including Kōbe (type locality), mainland China; "Korea"; Australia (Fain, 1974b; Fain, 1982; Zachvatkin, 1941; Okabe & Makino, 2002; Kawazoe at al., 2008). 1d02873439ae7d155a66708bb0d2e39e taxon_sennertia_alfkeni http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia alfkeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Alloxylocopa) appendiculata Smith, 1852 (=Xylocopa appendiculata circumvolans) (type host). ; Xylocopa (Alloxylocopa) amamensis Sonan, 1934; Xylocopa (Alloxylocopa) flavifrons Matsumura, 1912; Xylocopa (Alloxylocopa) albinotum Matsumura, 1926 201160b584083dad9948a7d839b0e886 taxon_sennertia_alfkeni http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia alfkeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This species was known as two separate species Sennertia alfkeni and S. japonica, but it was later shown that these two are genetically the same (Kawazoe et al., 2008) and morphological differences between them are probably continuous and size-dependant (Zachvatkin, 1941; Okabe and Makino, 2008).
Phoretic deutonymphs of the alfkeni-type attach themselves to the dorsoalateral hairs, while those of the japonica-type were common in the mesosomal and metasomal acarinaria of the host. The deutonymphs of both groups display allometric differences. The alfkeni-type deutonymphs are larger (311.8±36.9 μm), with the claw measuring 50.5±8.2 μm, and the ambulacra with a projection, whereas japonica-type deutonymphs are smaller (232.0±18.5 μm), with the claw size 14.7±2.2 μm, and lacking ambulacral projection (Okabe and Makino, 2002; Okabe et al., 2008; Kawazoe et al., 2008). The ambulacral projection found in alfkeni-type deutonymphs apparently aids in the attachment to the host’s hair. In contrast, deutonymphs of the japonica-type do not attach to hairs and are phoretic inside acarinaria, specialized internal structures on the host body serving for mite transfer.
Adults also expressed variation in the form of the dorsal setae, ranging from narrow to broad (Okabe et al., 2008). e497705bf4443615deedcab3912cb7f1 taxon_sennertia_americana http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia americana dispersing on the carpenter bee, Xylocopa virginica virginica, Illinois (BMOC BMOC 04-1222-159). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_americana_on_host_BMOC-04-1222-159.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8fd7755128d74f80bb46d68e651930ba taxon_sennertia_americana http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia americana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 distinctly longer than d1-h1; long, nearly as long as se; situated anterior to hysterosomal shield. Setae d1 and e1 nearly uniform in length with h1. Setae d1 shorter than 1/4 of distance between them; situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2; not touching hysterosomal shield. Lateral gland openings situated outside hysterosomal shield. Setae 4b, g, and 4a without distinct rhomb-like widening, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Posterior processes of coxal apodemes IV non-applicable. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple, mG II almost as long as leg II or longer. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, filiform. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical, not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV distinctly protruding apex of tarsus IV.
Adults. Supracoxal seta scx situated on supracoxal sclerite, anterior to outer ridge of supracoxal sclerite. Alveoli of setae ve absent. Dorsal idiosomal cuticle tuberculate or mammillate (except for posterio-medial opisthosomal region in female). Dorsal cuticular pattern more or less uniform. Dorsal idiosomal setae c1-h1 spiniform, short (not reaching half of distance to next posterior pair of setae). Dorsal idiosomal setae cp, c3, h3 widened distally, compressed dorso-ventrally at apex, barbs more numerous at apex. Dorsal setae e2 and f2 spiniform, distinctly shorter than h3. Prodorsal shield distinctly elongated, length/width 1.7-1.8, without falsifoveate pattern. Coxal fields III opened. Proximal acetabular extensions of ap' I partially border antiaxial margins of coxal fields I. Proximal acetabular extensions of ap' II partially border antiaxial margins of coxal fields II. Distal acetabular extensions of ap' II and ap'' II separate. Proximal acetabular extensions of ap' III completely border antiaxial margins of coxal fields III. Distal acetabular extensions of ap' III and ap'' III fused. Proximal acetabular extensions of ap' IV completely border antiaxial margins of coxal fields IV. Distal acetabular extensions of ap' IV and ap'' IV separate or not developed. Opisthosomal gland openings approximately at level of e2. Tarsal setae ra and la II present. Solenidion ω2 I subapical. Famulus ε spiniform. Setae ba I longer than famulus ε. Setae ba II absent.
Female. Setae ad1 and ad2 absent. Setae ps3 short, distinctly shorter than ps2; anterior to 4a level. External copulatory tube absent. Setae h3 nearly at level of h2. Posterio-medial part of dorsal opisthosoma with distinct longitudinal linear pattern.
Male. Setae ad1 absent. Genital setae short, transparent mammillae; situated on progenital folds. Pseudanal setae ps3 outside progenital sclerites, spiniform. Dorsal supporting sclerites short, as long as 2 diameters of aedeagus at base or shorter. Setae q I present, p II absent. Pretarsal suckers IV same as pretarsal suckers I-III.
Protonymph. Tarsal setae e IV absent, f IV absent.
Larva. Dorsal idiosomal setae relatively longer than in other instars, protruding bases of subsequent setae.
Notes. The slide marked "holotype" contains six specimens none of which is identified as the holotype. 070fbf729e08cdf4b97d7b0295ae1c39 taxon_sennertia_americana http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia americana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Florida, Illinois, Maryland, Michigan, New York (Albany Co., Albany - type locality), North Carolina, Pennsylvania, Texas, Virginia. c54d6cdbfef5b27ce6c5adf3bd08f7e5 taxon_sennertia_americana http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia americana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Xylocopoides) virginica (type host), Xylocopa (Xylocopoides) virginica texana. a5f0529723471b7aea5ec4977b8032da taxon_sennertia_americana http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia americana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Feeding instars of this species were found only once (Lombert et al. 1987). At the time these collections were made in early August, adult Xylocopa virginica were emerging from the nests, and most cells were already empty. A few cells contained pharate or teneral adult bees. Some cells contained large provision masses consisting of nectar and pollen, but no developing bee larvae. All instars of Sennertia americana were located on the walls of these cells. The cells were also inhabited by Horstia virginica Baker, 1962 (Acaridae) and Tortonia quadridens Baker, 1962 (Suidasiidae). Since the two latter mite species are cleptoparasitic the ultimate cause of cell failure is uncertain. abc392b8dc8df18c6a384bdc37d5413d taxon_sennertia_antarctica http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia antarctica, heteromorphic deutonymph (from Trägårdh, 1907). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_antarctica_orig.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9fb4bbcacff3259636ce01a5f168e5b9 taxon_sennertia_antarctica http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia antarctica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. The original description (Trägårdh, 1907) provides a brief description of phoretic deutonymph (number of studied specimens not specified, presumably one). The author also speculates that "Trichotarsus antarcticus " may be the deutonymphal stage of supralittoral mites of the genus Hyadesia. This insufficiently described taxon is similar to several African species belonging to the cerambycina-group of the genus Sennertia. The geographic locality is, most likely, erroneous because carpenter bees, the only hosts of Sennertia, do not occur in Antarctica efcdb8057096d7532c61fa1df72c94cc taxon_sennertia_antarctica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia antarctica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution "Antrarctica, Booth Is. (as Wandel Is.)" (Trägårdh, 1907) 4e13afa39d3ac0056accd0d8424764f5 taxon_sennertia_antarctica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia antarctica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "marine algae" (Trägårdh, 1907) 74fd4b0c2c8dfc096593d3e657ae9f6b taxon_sennertia_argentina http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of the mite Sennertia argentina, ex Xylocopa mexicanorum, Mexico (BMOC 05-0102-027). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_argentina_BMOC_05_0102_027.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3b5a1e32979973925cb9ec0a71adf050 taxon_sennertia_argentina http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymph of the mite Sennertia argentiana, emerging from the genital acarinarium of the bee Xylocopa frontalis, Panama (BMOC 04-1222-157). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_argentina_on_host_BMOC-04-1222-157f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ec134e5479f4ad9002c4c64dff3ab3e4 taxon_sennertia_argentina http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia argentina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part distinctly narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 and d1-h1 uniform in length, microsetae. Setae c1 microsetae, situated anterior to hysterosomal shield. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2; not touching hysterosomal shield. Setae h3 shorter or nearly equal to legs IV. Lateral gland openings situated outside hysterosomal shield. Setae 1a, 3a, 4a attenuated. Setae 4b, g, and 4a without distinct rhomb-like widening but widened otherwise (spiniform or nearly spiniform). Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II spiniform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, arc-like. Posterior apodeme IV present, not connected to anterior apodeme III. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 enlarged. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple, mG II distinctly shorter than leg II, but longer than femur II. Tarsal setae la I-II microsetae, as long as famulus ε. Tarsal setae ra I-II not bifid, spiniform. Tarsal setae wa I-II and s III spiniform (at least s III with rounded apices). Tarsal setae d I-II distinctly widened, lanceolate. Tarsal setae d and f I-II strongly asymmetrical, f about 2 times shorter and filiform, e longer and lanceolate; d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore absent. Anterior condylophore I-II without distal bending. Seta d III situated shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV shorter than femur IV.
Other instars unknown. a5e8294b90253d44610fdfb742f2cfbf taxon_sennertia_argentina http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia argentina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina (type locality); Brazil; Costa Rica; Guatemala; Mexico: Jalisco, Sinaloa, Chiapas; Nicaragua; Peru; Panama; Suriname. bb10a17b87326a4c644da23c332564b4 taxon_sennertia_argentina http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia argentina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) frontalis (type host), Xylocopa (Neoxylocopa) fimbriata, Xylocopa (Neoxylocopa) nautlana, Xylocopa (Neoxylocopa) nasica, Xylocopa (Neoxylocopa) mexicanorum (Apidae). 9bc8b155b9e96367b71e85d421a50d19 taxon_sennertia_argentina http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia argentina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Frequently found phoretic in and around genital capsules of host males, numerous deutonymphs were also found in a special pouch (genital acarinarium) of the female genital system (Klimov et al., 2007b). 88cbb6bd7a73483dab9e49e2a483633c taxon_sennertia_augustii http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia augustii, heteromorphic deutonymph, Argentina, ex Xylocopa augusti. http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_augustii_BMOC_04-1112-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 27f0199fe0455dabd8e075c7414cc728 taxon_sennertia_augustii http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia augustii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. This taxon can not be separated without an overlap from Sennertia frontalis and, especially Sennertia shimanukii in univariate and multivariate morphometric space (Klimov & OConnor, 2008). A molecular analysis is necessary to validate these taxa. 792b9f7a5a0918cccfe99581cb8a02ea taxon_sennertia_augustii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia augustii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina: Buenos Aires (Berazategui) (type locality), Uruguay, (?) Mexico 6bfc4c67e7ea6e003e83a3a40c733541 taxon_sennertia_augustii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia augustii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) augusti, (?) Xylocopa fimbriata. c44f7d697d082d54f4d582f225677355 taxon_sennertia_basilewskyi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia basilewskyi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Haut-Congo Prov. ("Uélé") (Bambesa) (type locality), Dingila. 4f22cf5421dd667dce66b209b3a13a33 taxon_sennertia_basilewskyi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia basilewskyi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "Megachilidae MRAC n° M 42" (Bambesa) (type host), "Chedrion nigrihirtum" (Dingila), Xylocopa (Koptortosoma) africana (as Mesotrichia africana) (Bambesa) (Fain, 1974; Fain, 1981) ee9461d478dd5e5a39cb6ea8aec7f19c taxon_sennertia_benoiti http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia benoiti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Côte dʼIvoir: Bafing, 7 km ESE Touba (type locality), Democratic Republic of the Congo 1ba9c07c451428dc86de1f655d43009d taxon_sennertia_benoiti http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia benoiti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "Ceratina sp. (n° 283 A) (Fain, 1974a) or (n° 283 B) (Fain, 1980), Ceratina (Pithitis) atopura (Democratic Republic of the Congo) 01b63399e674a33b151d54477367d32c taxon_sennertia_bifida http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia bifida, heteromorphic deutonymph, ex Ceratina flavipes, Japan (BMOC 95-0422-046). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_bifida_BMOC_95-0422-046.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2d7bb0067e1a4c1a66eb1d0fef2c4e93 taxon_sennertia_bifida http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia bifida http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan: Hokkaido (Sapporo, Hitsujigaoka) (type locality), Honshu, Ryukyus 5b3b62b3200ea262df837f044f24f756 taxon_sennertia_bifida http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia bifida http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Ceratinidia) japonica Cockerell, 1911(type host); Ceratina flavipes Smith, 1879; Ceratina okinawana Matsumura and Uchida, 1926 618cfabb9c7472929854a45d42930714 taxon_sennertia_bifilis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Sennertia bifilis (larger mites) and Horstia trifilis (smaller) on Xylocopa combinata  (BMOC-05-0420-115). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_bifilis_Horstia_trifilis_on_hostBMOC-05-0420-115.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 29ed115124bad5bf0c569528f98f18aa taxon_sennertia_bifilis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Sennertia bifilis (larger mites) and Horstia trifilis (smaller) on Xylocopa combinata  (BMOC-05-0420-115). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_bifilis_Horstia_trifilis_on_hostBMOC-05-0420-115b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 27bd74b7c6d25836c085293d621e285e taxon_sennertia_bifilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia bifilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Papua New Guinea: Madang: Astrolabe Bay (Erima) f6b22d21a1c1de68da6802ecfd5e01af taxon_sennertia_bifilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia bifilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) combinata Ritsema, 1876 9073a87376aa7f65c570fffb75d70ce1 taxon_sennertia_caffra http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia caffra http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution South Africa: Eastern Cape (Willowmore) ("Willowmore, Kapland") 15f720ace3f20d36d5f92759d528d0f8 taxon_sennertia_caffra http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia caffra http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) caffra (as Koptorthosoma caffra) 1e1b3740c2340ae1ff4f88cfde328c96 taxon_sennertia_cantabrica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia cantabrica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Algeria 1df2e143d14b297bb7088a7fa6419062 taxon_sennertia_cantabrica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia cantabrica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Xylocopa) cantabrita Lepeletier, 1841 28e4c327658d6374f63eeda128ce3cc7 taxon_sennertia_capensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia capensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution South Africa: Eastern Cape (Willowmore) 3e1e49e4d6ddc4e65d39c34ce382393b taxon_sennertia_capensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia capensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Gnathoxylocopa) sicheli Vachal, 1898 86a6e7c852bdc196cd33275bdafe80f2 taxon_sennertia_cerambycina http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia cerambycina, heteromorphic deutonymph, ex Xylocopa violacea, France (BMOC 05-0420-424). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_cerambycina_BMOC_05-0420-424.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7b5fe1e5f28763dadbfe0bbbe07721cb taxon_sennertia_cerambycina http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia cerambycina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution "Carniola" (in modern Slovenia) (type locality), Greece, France, Italy, Spain, Germany, Netherlands, Iran, China (Donnadieu, 1868; Canestrini & Fanzago, 1878; Eyndhoven, 1941; Scopoli, 1763; Vicidomini & Meloni, 1999; Vitzthum, 1912c; Zachvatkin, 1941; Fain, 1981a), "All Europe" (Berlese, 1897), "All southern, central, and eastern Europe, up to 55°N; West and Central Asia up to Tibet and Gashun Gobi Desert (China) on the east and up to northern Iran to the south" (Zachvatkin, 1941). 5d2f84b6ced728266617fbf0c1d01c2e taxon_sennertia_cerambycina http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia cerambycina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa violacea (Linnaeus, 1758)(as Apis Violaceae) (type host),; Xylocopa (Xylocopa) valga Gerstäcker, 1872; Xylocopa (Xylocopa) varentzowi Morawitz, 1895; Xylocopa (Proxylocopa) nitidiventris Smith, 1878; Xylocopa (Proxylocopa) przewalskyi Morawitz, 1886; Xylocopa (Copoxyla) turanica Morawitz, 1875; Xylocopa (Copoxyla) iris (Christ, 1791); Polochrum repandum Spinola, 1806 (Hymenoptera, Sapygidae) (cleptoparasite of Xylocopa); (Scopoli, 1763; Vitzthum, 1912c; Zachvatkin, 1941; Vicidomini, 2005); ; Occasionally found phoretic on the following bee hosts:; Hoplitis princeps (Morawitz, 1872) (as Osmia princeps); Osmia (Monosmia) apicata Smith, 1853(as Osmia macroglossa); Megachile (Eutricharaea) leachella Curtis, 1828 (as Megachile argentata); Anthophora (Paramegilla) balassogloi (Radoszkowski, 1876) (as Anthidium christofi F. Mor.); (Zachvatkin, 1941) e16b77aec11f7e6cf6f3bc16fdc74c5e taxon_sennertia_cerambycina http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia cerambycina http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Normally associated with Xylocopa (see below), feeding on the provisioned pollen in the bee nest. Rarely dead bee eggs and larvae can be found in the infested cells. However, it is unclear if the mite presence causes the death or the mite can kill the bee eggs or larvae directly (Vicidomini, 1996).
Adult bees of Xylocopa violacea extensively infested with S. cerambycina show slow flight, lack of precision in flying trajectories, difficult and imprecise landing, heavy take-off, and reduced feeding rates (Vicidomini, 1999). Phoretic mites may block the epinotal spiracles, causing asphyxia in the bee host (Vicidomini, 1999).
Duchemin (1886) reports death of 30 honeybee hives in France attributable to this mite. The mite infests honeybees visiting sunflowers, eventually causing death of colonies. Similar cases (e. g., death of honeybee colonies with the introduction of sunflower) were reported in Russia at the end of 19th - beginning of 20th century (reviewed Grobov, 1978). Experimental data suggesting that the mite is responsible for the death of bees are, however, absent. 2c3d63256fbb50af04f3e61164768fcf taxon_sennertia_ceratinarum http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia ceratinarum, heteromorphic deutonymph, ex "Ceratina pilipes", Mozambique (BMOC 04-0508-288). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_ceratinarum_BMOC_04-0508-288.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b35fb36d85dee2d6ae1da86c076211a4 taxon_sennertia_ceratinarum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia ceratinarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution South Africa: KwaZulu-Natal (Salt Rock) (type locality); Democratic Republic of the Congo; Cameroon (Fain, 1974a, 1980, 1981a); Mozambique (our data, unpubl.) b363c3567abef656ce2f297a07a346ce taxon_sennertia_ceratinarum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia ceratinarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Pithitis) sp. (type host) (Fain, 1981a) (note: type host was also indicated as Ceratina inermis Friese, 1905 (as Ceratina (Pithitis) turneri (Fain, 1974a)) and "Ceratina nr. turneri" (Fain, 1980))); Ceratina nasalis Friese, 1905 (as Ceratina acutipyga) (South Africa); Ceratina atopura Cockerell, 1937 (Congo); Ceratina (Pithitis) nilotica Cockerell, 1937 (Congo); Ceratina (Pithitis) viridis Guérin-Méneville, 1844 (as Ceratina congoensis and Pithitis viridis) (Cameroon, Congo); (Fain, 1974a, 1980, 1981a); "Ceratina pilipes" (our data, unpubl., host name not verified) c1b6b380e3dc86fb6c534b90bbc90692 taxon_sennertia_congoicola http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia congoicola http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo 3b38ec90d72c981a06878edef298e046 taxon_sennertia_congoicola http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia congoicola http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "Mesotrichia striata" (type host) ; Xylocopa (Koptortosoma) imitator Smith, 1854; Xylocopa (Mesotrichia) torrida (Westwood, 1838); (Fain, 1971, 1981a) dbc03e612cb72107f425bee41a9ab2a7 taxon_sennertia_dalyi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia dalyi, heteromorphic deutonymph, ex Ceratina chrysomalla, Cyprus (BMOC 04-0508-277). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_dalyi_BMOC_04-0508-277.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 05b04de38e61bea74aa3317340f306c0 taxon_sennertia_dalyi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia dalyi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tunisia: Jundūbah, 3.5 S Tabarka (type locality), ‘Ayn ad Darāhim; Silyānah (Maktar); Nābul (Grombalia) (Fain, 1980, 1981a); Cyprus (our data, unpubl.). a9dc6a98f2ce475307ed76c7725f6f14 taxon_sennertia_dalyi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia dalyi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Euceratina) dallatorreana Friese, 1896 (type host); Ceratina (Euceratina) chalybea Chevrier, 1872 (Maktar); Ceratina (Euceratina) mocsaryi Friese, 1896 (Ain Draham); Ceratina (Euceratina) callosa (Fabricius, 1794) (Grombalia); (Fain, 1980, 1981a); Ceratina (Euceratina) chrysomalla Gerstäcker, 1869 (Cyprus) (our data, unpubl.) c93cfe05da8ab0d55175278938a9628b taxon_sennertia_dalyi_nilotica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia dalyi nilotica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Egypt: Al Qalyūbīyah, Banhā ("Benha, Delta du Nil") (type locality) 6a643e57054610c143ad00744a86c9bc taxon_sennertia_dalyi_nilotica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia dalyi nilotica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "Pithitis sp. n. A" (type host); Ceratina (Pithitis) tarsata Morawitz, 1872 (as Pithitis tarsata); "Ceratina sp. n. " (Fain, 1980) 9a896b1f41f91ded62baf49a6105d2d6 taxon_sennertia_delfinadoae http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia delfinadoae, holotype, phoretic deutonymph, ex Platacanthomys lasiurus (Rodentia, Platacanthomyidae) (accidental host), India, BMOC 11-0930-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_delfinadoae_holotype_HDN_BMOC_11-0930-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 83f0cda1293f5adba880ad548381e161 taxon_sennertia_delfinadoae http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia delfinadoae, holotype slide, IRSNB, BMOC 11-0930-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_delfinadoae_Holotype_slide_IRSNB_BMOC_11-0930-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 668dd86342b806dc8928b0bb00165677 taxon_sennertia_delfinadoae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia delfinadoae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution India e2a4c8ad7a7a7957a9b77730c62c0c4f taxon_sennertia_delfinadoae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia delfinadoae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "rat" (type host, acknowledged as accidental) (Fain, 1981a); Xylocopa (Nodula) amethystina (Fabricius, 1793) 7b74c2e36de65d000a195566db900445 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia devincta, phoretic deutonymph, ex Ceratina sp., Peru (BMOC 03-0604-016). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_devincta_BMOC_03-0604-016_holotype.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8396914921f256b382933feef4e51fd2 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia devincta near the entrance of the metasomal acarinarium of the bee, Ceratina sp., Peru (BMOC 03-0604-016). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_devincta_nr_acarinarium_BMOC_03-0604-016.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2b167265943d6eb0e515485eb580717f taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia devincta dispersing on the bee, Ceratina sp., Peru (BMOC 03-0604-016). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_devincta_on_host_BMOC_03-0604-016.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7ff633e37223b418e9de477a4d1ac937 taxon_ http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Deutonymph. Gnathosomal solenidia distinctly shorter than 1/3 width of femur I. Supracoxal setae scx situated on separate small sclerites. Hysterosomal shield distinctly expanded beyond opisthonotal gland openings and bases of setae f2. Lateral edges of hysterosomal shield in anterior part not narrowing, rounded. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeding diameter of si alveoli. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization; non-uniform at posterior end of hysterosomal shield, with two distinct areas. Striae long. Distinct vestiges of vi present, situated at level of se. Setae si distinctly posterior to se. Setae si slightly longer than se, as thick as se. Setae c1 long, nearly as long as se, distinctly longer than d1-h1, situated on anterior edge of hysterosomal shield. Setae c3 nearly spiniform, situated on small, weakly developed sclerite. Setae cp extending beyond bases of e2. Setae d1 and e1 similar in length to h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2. Setae e2 not touching hysterosomal shield. Lateral opisthonotal gland openings situated on hysterosomal shield. Setae 1a, 4b, and 3a conoidal; g and 4a spiniform, with attenuated tips. Posterior apodemes II and anterior apodemes III partially fused. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV present, not connected to anterior apodeme III. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+ad2). Conoids ps2 anterior to ps1, situated outside outer level of ad1+ ad2. Suckers ad3 not enlarged, smaller than central suckers, without rough sclerotization. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present, very small. Posterio-medial longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Legs with setae pR I-II, sR III, wF IV, gT I-II, kT III filiform. Genual setae mG I-II simple, shorter than leg II, but longer than femur II. Tarsal setae la I-II microsetae. Tarsal setae ra I-II not bifid, spiniform. Tarsal setae wa I-II spiniform, bifid apically, s III spiniform, simple. Tarsal setae d I-II foliate, shorter than f I-II. Alveoli of tarsal setae d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II bent distally. Setae sR III extending beyond femur III. Seta d III apically shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV extending beyond posterior edge of hysterosoma. Tarsus IV not enlarged, shorter than width of trochanter IV. Setae w IV almost half as thick as d IV, distinctly longer than leg IV. Setae w IV situated on middle of tarsus IV. Setae s IV present, very short. Setae wF IV not extending beyond tibia IV. e32c5b01a2550f01df42fd92afc344f8 taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Peru (type locality). ed25b6e64311183d6934c745845fb1d4 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina sp. 54fc55cd412e8862f557b8e8cd968a58 taxon_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the metasomal acarinarium of the host. 4fce4d623cb7927e24dfcb562595cf10 taxon_sennertia_dissimilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia dissimilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan (Zachvatkin, 1941). The bee host was probably originated from a Southeast or East Asian country occupied by Japan at the collection time (Okabe et al., 2010). 90c55ee7f41aca4623d5a78a61280c2f taxon_sennertia_dissimilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia dissimilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Biluna) nasalis Westwood, 1842 (as Xylocopa dissimilis) (type host) 5fd551bc48fdb222ce150611370743e6 taxon_sennertia_donaldi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia donaldi, phoretic deutonymph, ex Xylocopa frontalis, Brazil, HK 84-0820-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_donaldi_HK_84-0820-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3459d6894b24ff32d158ea591c20badd taxon_sennertia_donaldi http://purl.org/dc/dcmitype/StillImage image/jpeg syntype slide http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_donaldi_syntypes.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 6a244675e5d7a23c49ccdda30e0557f1 taxon_sennertia_donaldi http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia donaldi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Similar to Sennertia argentina, see differences in the key to species of Sennertia.
Notes. Fain (1981a) synonymized this species with Sennertia argentina. We found distinct differences between S. argentina and S. donaldi (see key) and, therefore, consider the latter as a valid species. 257e2340ce2ee4d0f1bd916e3d70a060 taxon_sennertia_donaldi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia donaldi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Trinidad and Tobago (type locality), Brazil, Venezuela. 225648a6059361a3561d7e4f060dd602 taxon_sennertia_donaldi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia donaldi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) frontalis (Olivier, 1789) (type host); Xylocopa (Neoxylocopa) fimbriata Fabricius, 1804 426aa8cca53b3eb0856fb6ff269e7182 taxon_sennertia_duweinii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia duweinii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Originally not indicated, probably Egypt. 947f2cf70f0489c3625d775c311bc863 taxon_sennertia_duweinii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia duweinii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) aestuans (Linnaeus, 1758) 78868c0fab44f4102ba8659af9d9e078 taxon_sennertia_duweinii http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia duweinii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Deutonymphs were collected inside acarinarium (Sherbef & Duweini, 1980). The type of acarinarium is not specified, but probably it is the metasoma acarinarium. 0e2ca580a4265c0b5669e969cbad2c54 taxon_sennertia_egyptiaca http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia egyptiaca http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Egypt: Al Qāhirah, Shubrā al Khaymah ("Shoubra Elkheima, Cairo") (type locality); Sudan 51c8189691926330802f2f5ae3cc014c taxon_sennertia_egyptiaca http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia egyptiaca http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) aestuans 32c293f5c1f7efa6367c743e0accea8a taxon_sennertia_egyptiaca http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia egyptiaca http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the metasomal acarinarium (Elbadry, 1971). 2a61780898df1d5b64b7e9809472a082 taxon_sennertia_elseni http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia elseni, phoretic deutonymph (HK 85-0117-002) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_elseni_HK_85-0117-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7ead36220f865fddcf6ae5561e4573b2 taxon_sennertia_elseni http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia elseni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Bas-Congo (Muanda) (as "Moanda") (type locality); Madagascar (Fain, 1971; Fain & Pauly, 2001). 9c4b8d408e17930a49bf235330b3fef0 taxon_sennertia_elseni http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia elseni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) caffra (Linnaeus, 1767) (as Mesotrichia olivacea) (type host); Xylocopa (Koptortosoma) calens Lepeletier, 1841 (Madagascar); Melanempis sp. (Madagascar) (cleptoparasitic bee of the subfamily Nomadinae); (Fain, 1971; Fain & Pauly, 2001) 143563ad2e0e54adad0b3e46609771f2 taxon_sennertia_faini http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia faini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Similar to Sennertia lucrosa (see notes on Sennertia lucrosa and key to species). eca4dcd7f6c768b6c142648600c0b628 taxon_sennertia_faini http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia faini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Guatemala: Sololá 3087971b2c6cba3795177be523602c37 taxon_sennertia_faini http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia faini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Apis mellifera (alcoholic washings of dead honeybees and on bee hairs) 7e7175dc0d1971cf5c32e655ca2e6643 taxon_sennertia_flabellifera http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia flabellifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java (Bogor) (as "Buitenzorg") 8057f91f98cb4dea7ae8dc5c354eca8f taxon_sennertia_flabellifera http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia flabellifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) sp. (as Koptorthosoma sp.) 3f3c92d48cf25f8515cc20d7a4e2e9a1 taxon_sennertia_frontalis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia frontalis dispersing on the bee, Xylocopa frontalis, Venezuela, BMOC 04-1122-026 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia-frontalis_on_host_BMOC_04-1122-026.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 0b1c977c31c89f11cec5d651713265b8 taxon_sennertia_frontalis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia frontalis, ex Xylocopa frontalis, Venezuela, BMOC 05-0420-204 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_frontalis_BMOC_05-0420-204.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 157e72ed18552d0bc0d13a2be59bd562 taxon_sennertia_frontalis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia frontalis dispersing on the bee, Xylocopa frontalis, Venezuela (BMOC 05-0420-204). http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia-frontalis_on_host_BMOC_04-1122-026b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b359b97c4b66316a511fed94daa82565 taxon_sennertia_frontalis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia frontalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Similar to Sennertia augustii and S. shimanukii (see Klimov & OConnor, 2008).
Other instars unknown. 580eae2a3870a9ee9397e41477454a7b taxon_sennertia_frontalis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia frontalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina (type locality): Entre Ríos; Brazil: Amazonas, São Paulo; Suriname: Marowijne; Peru: Loreto; Venezuela: Monagas; Panama: "Canal Zone"; Honduras: Progresso; Mexico: Chiapas 60f4a0137b602bda8c449d375dcbfae6 taxon_sennertia_frontalis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia frontalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) frontalis (type host), Xylocopa (Neoxylocopa) nautlana. a05e61038c40c0ee31e9d0627137181d taxon_sennertia_gargantua http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia gargantua http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Uzbekistan, Tajikistan (Zachvatkin, 1941), Kyrgyzstan (Grobov, 1975), Iran (Klimov, unpublished) 71ac68e08c5bfe6293ef70a5a0bd355d taxon_sennertia_gargantua http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia gargantua http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Nodula) punctilabris Morawitz, 1894 ; Xylocopa (Xylocopa) valga Gerstäcker, 1872; (Zachvatkin, 1941) 5c1d07f31813d78f7cc31b0456492377 taxon_sennertia_greeni http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia greeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Sri Lanka: Central (Eton estate) ("Eton Estate, Punduloga, Ceylon") e3070d27b7403a58a93b7775bc0417d2 taxon_sennertia_greeni http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia greeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Found in nest of Xylocopa (Mesotrichia) tenuiscapa Westwood, 1840 (as Koptorthosoma tenuiscapa). 29bb455e0869f63319bcfe1818ac2c46 taxon_sennertia_haustrifera http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia haustrifera, phoretic deutonymph, holotype, ex Ceratina capitosa, Mexico, BMOC 97-0331-035 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_haustrifera_holotype_BMOC_97-0331-035.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 46ffea1d336a7da9b79e41fe910f368e taxon_sennertia_haustrifera http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia haustrifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on dorsal extension of posterior apodeme I. Hysterosomal shield distinctly expanded beyond lateral gland opening and bases of setae f2. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield formed by long striae, with sclerotization, sclerotization shifted to posterior striae. Distinct rudiments of vi absent. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 and d1-h1 uniform in length, microsetae. Setae c1 microsetae; situated posterior to anterior margin of hysterosomal shield. Setae c3 conoidal, situated on large triangle sclerite touching posterior apodemes II and anterior apodemes III. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated on hysterosomal shield. Setae 4b conoidal. Setae 4b, g, and 4a without distinct rhomb-like widening (4b pear-shaped, 4a and g filiform). Setae pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV present, connected to anterior apodeme III. Conoids ps2 anterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) with rough sclerotization, maximal length of rough sclerotization distinctly shorter than two diameters of anterior suckers. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers absent. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II bifid at tips, mG II shorter than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, filiform. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II foliate. Tarsal setae d and f I-II almost symmetrical, d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV not protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV not protruding tibia IV.
Other instars unknown. 2f695e8aa1cb09dd390788f9e4311a4c taxon_sennertia_haustrifera http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia haustrifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Jalisco (type locality), Chiapas. e4ecfe77d996ec58074dd131fa782f53 taxon_sennertia_haustrifera http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia haustrifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina capitosa Smith, 1879 (Apidae). d7717efb60f19e0f0745ef8556ec3072 taxon_sennertia_herminae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia herminae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Madagascar: Antananarivo (Antananarivo) 72b0a376e7871cd7f4cdd89a21d54ebb taxon_sennertia_herminae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia herminae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa sp. 2bbadf6a1ed6b2c47be0f2b5fda5e356 taxon_sennertia_hipposideros http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia hipposideros, phoretic deutonymph (HK 85-0120-5) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_hipposideros_HK_85-0120-5.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e eb5405e6c6dd6ff020e1444f23eee7af taxon_sennertia_hipposideros http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia hipposideros http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution India (type locality), Malaysia (Oudemans, 1902; Fain, 1981a), Indonesia, Philippines c7c78775f6b86652900c81f8174ab1ff taxon_sennertia_hipposideros http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia hipposideros http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) tenuiscapa Westwood, 1840 (type host) (India); Xylocopa (Mesotrichia) latipes (Drury, 1773) (Malaysia, Indonesia, Philippines) 4fab5cfab299fd56cbc29ff43797de08 taxon_sennertia_hipposideros http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia hipposideros http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Phoretic inside the metasomal acarinarium of the host (Oudemans, 1902; OConnor, 1993b). 33e8afde996fbbdfacbb36e730f4672a taxon_sennertia_horrida http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia horrida dispersing on the bee, Xylocopa mcgregori, Philippines, BMOC 05-0420-055 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_horrida_BMOC_05-0420-055.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 481d664bf7122a44181d2f12713f85a0 taxon_sennertia_horrida http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia horrida dispersing on the bee Xylocopa iridipennis, Indonesia (BMOC 05-0420-052). Click to enlarge, BMOC 05-0420-052 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_horrida_on_host_BMOC-05-0420-052.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e caf8218eb34a8014d1a5ff6abca42972 taxon_sennertia_horrida http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia horrida dispersing on the bee Xylocopa iridipennis, Indonesia (BMOC 05-0420-052). Click to enlarge http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_horrida_on_host_BMOC-05-0420-052c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c818e33b12b330c78f8956577bf781b9 taxon_sennertia_horrida http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia horrida http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java, Jakarta (as "Batavia") (type locality) (Vitzthum, 1912d), Sumatra (Oudemans, 1924); Japan; India: Assam (Zachvatkin, 1941), Tamil Nādu (Ramaraju & Mohanasundaram, 2001), Phillipines (Klimov & OConnror, 2008) 966b9ea7841b76c799eeefd7c237f732 taxon_sennertia_horrida http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia horrida http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Biluna) nasalis Westwood, 1842 (as Xylocopa (Koptorthosoma?) dissimilis) (type host); Xylocopa (Koptortosoma) aestuans (Linnaeus, 1758); Xylocopa auripennis Lepeletier, 1841 ; Xylocopa fallax Maidl, 1912 ; Xylocopa iridipennis Lepeletier, 1841 ; Xylocopa mcgregori Cockerell, 1920 ; Xylocopa mimetica Cockerell, 1915; (reviewed Klimov & OConnror, 2008) c88aad147d45bc37cda3e2ade2e556a3 taxon_sennertia_hurdi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia hurdi, phoretic deutonymph, ex Xylocopa tabaniformis tabaniformis, Mexico, BMOC 04-1122-021 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_hurdi_BMOC_04-1122-021.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 0514d2c7238f9590cd62226257d925c9 taxon_sennertia_hurdi http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia hurdi dispersing on the carpenter bee, Xylocopa tabaniformis azteca, Mexico, BMOC 04-1122-023 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_hurdi_on_host_BMOC-04-1122-023a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e f87f2f0e690dee9a168883ea8acd23d2 taxon_sennertia_hurdi http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia hurdi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph . Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 almost 2 times longer than d1; distinctly longer than h1; c1 long, nearly as long as se; situated anterior to hysterosomal shield. Setae d1 and e1 distinctly longer than h1; situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2; not touching hysterosomal shield. Setae h3 shorter than 110. Lateral gland openings situated outside hysterosomal shield, with usual distinct transverse slit. Setae 4b, g, and 4a without distinct rhomb-like widening, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Setae 4a not reaching transverse level of pores ip. Posterior apodemes II and anterior apodemes III free. Posterior processes of coxal apodemes IV not reaching level of anterior cuticular suckers rudiments. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, short. Ventral hysterosoma smooth. Genual setae mG I-II simple; mG II almost as long as leg II or longer. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, blade-like. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical; d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Seta wF IV almost reaching apex of tarsus or slightly protruding it.
Other instars unknown.
Notes. Closely related to S. ignota, S. longipilis, and an undescribed species from X. funesta. The differences between them are given in the key to species. c3d0ce43515feff5b89e8f66133506eb taxon_sennertia_hurdi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia hurdi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Oaxaca. e9d4ef8990aed734539ad8652daac52a taxon_sennertia_hurdi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia hurdi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Notoxylocopa) tabaniformis tabaniformis; Xylocopa tabaniformis azteca. 1f4438a6353cc08017fe7471d8d6f98f taxon_sennertia_ignota http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia ignota, phoretic deutonymph, ex Xylocopa viridis , Costa Rica, BMOC 05-0102-022 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_ignota_BMOC_05-0102-022.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 03f43dbfc29e5ab38c2ebd374c3d1bc7 taxon_sennertia_ignota http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia ignota http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Description of HDN, 10 paratype HDNs mentioned originally (Delfinado & Baker, 1976) c23ac307667aef54ae2c26156167796f taxon_sennertia_ignota http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia ignota http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Peru: Piura (Talara) (type locality); Costa Rica (our data) c6729ac70515ec35aabe97c50546beea taxon_sennertia_ignota http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia ignota http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa sp. (type host); Xylocopa viridis Smith, 1854 (our data) 976145595bd51978a6d4df8db67155f3 taxon_sennertia_indica http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia indica, phoretic deutonymph, ex Ceratina smaragdula, China, BMOC 97-0331-041 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_indica_BMOC_97-0331-041.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 878a58d21005ee9599461bf357ee1836 taxon_sennertia_indica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia indica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution India: Haryāna (Hisār) (type locality, as Hissar) (Delfinado & Baker, 1976); China (our data) 486e4d723352b3f27259364d66f3912d taxon_sennertia_indica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia indica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina binghami Cockerell, 1908 (type host, as Tithitis binghami); Ceratina smaragdula (Fabricius, 1787) e6af1ffb88b372f95001fe70286b0a4a taxon_sennertia_jeanalexi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia jeanalexi, phoretic deutonymph, ex Xylocopa nigrita, Cameroon, BMOC 03-0630-007 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_jeanalexi_BMOC_03-0630-007.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9b6114656bdc3b6109bbbbbdb00375cc taxon_sennertia_jeanalexi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia jeanalexi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: "Bubulu river, near Mvungu, Mayumbe" (type locality), (Bambesa Fain, 1971); Cameroon (our data) de92e3db9ca2d176e86eafba5cdae6ca taxon_sennertia_jeanalexi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia jeanalexi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) nigrita (Fabricius, 1775) (type host from Bubulu) (Cameroon); Xylocopa (Koptortosoma) lepeletieri Enderlein, 1903 (Bambesa) ae66107eca3642379f0913a86033a24c taxon_sennertia_koptorthosomae http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia koptorthosomae, phoretic deutonymph, ex Xylocopa latipes , Indonesia (Sumatra), BMOC 90-1212-006 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_koptorthosomae_BMOC_90-1212-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3f406724217f60cce21175468e249ea2 taxon_sennertia_koptorthosomae http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia koptorthosomae sporotheca http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia-koptorthosomae_sporotheca_BMOC_90-1212-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e aacb7ca97aae2d10236dbd4c2c3ef04d taxon_sennertia_koptorthosomae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia koptorthosomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java (type locality), Sumatra ("Medan, Deli") (Fain, 1974b), Malaysia (OConnor 1993b) bf6bdc90a06ef90ba05d65ea224eedbf taxon_sennertia_koptorthosomae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia koptorthosomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) tenuiscapa Westwood, 1840 (type host); Xylocopa (Mesotrichia) latipes (Malaysia); (Fain, 1974b; OConnor, 1993b) fe500b9abc753019acc0d32792fb07f5 taxon_sennertia_koptorthosomae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia koptorthosomae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in metasomal acarinarium (OConnor, 1993b). Dorsal hysterosoma with a distinct, elongated sporotheca, serving for transport of fungal spores (Klimov & OConnor, 2008). The mite probably feeds on the fungi inside the bee nest. 2764dcebe08c36938001b2c96b6f0833 taxon_sennertia_latipilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia latipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Kenya: "Malindi" ca51b363cea252f0fd652b3ae211bf42 taxon_sennertia_latipilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia latipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "Apidae (n° 139A)" (type host); Ceratina (Ctenoceratina) penicilligera Strand, 1912 e34c89641cabb97fe09e617e43b58237 taxon_sennertia_lauta http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia lauta, phoretic deutonymph, ex Xylocopa fuliginata, Philippines, BMOC 05-0420-140 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_lauta_BMOC_05-0420-140.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 43d41a7c3c926a5f5c1672e0bb8ff0ac taxon_sennertia_lauta http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia lauta dispersing on the male carpenter bee, Xylocopa fuliginata, Philippines , BMOC 05-0420-210 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_lauta_on_host_BMOC_05-0420-210a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e ce0b56f696ed74cd4d07423a8325d368 taxon_sennertia_lauta http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia lauta dispersing on the female carpenter bee, Xylocopa fuliginata, Philippines , BMOC 05-0420-209 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_lauta_on_host_BMOC_05-0420-209.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8bc52359f7f136188f0356c77a6a20e5 taxon_sennertia_lauta http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia lauta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Deutonymph. Gnathosomal solenidia shorter than 1/3 width of femur I, longer than distance between them. Distance between gnathosomal solenidia distinctly exceeds diameter of alveolus of vF I. Rostral projection without sclerite. Supracoxal setae scx situated on separate small and rounded sclerites. Alveoli of vi anterior to level of se and si; distance between them shorter than distance si-si. Lateral edges of hysterosomal shield in anterior part not narrowing, rounded. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si substantially exceeds diameter of si alveoli. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization; formed by long striae. Distinct vestiges of vi present. Setae si slightly anterior to se; longer than 1/3 of se; almost as thick as se. Setae c1 and d1-h1 uniform in length, microsetae. Setae c1 microsetae; posterior to anterior margin of hysterosomal shield. Setae c3 distinctly widened. Setae d1 situated on hysterosomal shield. Cupules ia situated closer to alveoli of c2 than to alveoli of cp. Sclerite between ia and d2 absent. Setae e2 subequal with d2, width not exceeding diameter of their alveoli. Alveoli of setae e2 not touching hysterosomal shield. f2 situated lateral to hysterosomal shield. Lateral opisthonotal gland openings situated on hysterosomal shield. Cupules im situated approximately on line connecting d2-e2, closer to e2 than to d2. Setae 4b, g, and 4a widened but without distinct rhomboid widening. At least pR I-II, sR III, wa I-II, wF IV, gT I-II, hT I-II, and kT III filiform or nearly filiform. Coxal setae 1a inflated at basally. Coxal setae 4b widened at base, alveolar sclerites of 4b, if present, not fused to posterior apodemes II. Alveolar sclerites of c3 and 3a present or absent not fused with anterior apodemes III. Posterior apodemes II and anterior apodemes III distinctly fused. Anterior portion of posterior apodeme II distinct, well-sclerotized before its fusion to anterior apodeme III. Anterior apodemes IV arc-like, not interrupted.. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+ad2). Conoids ps2 anterior to ps1, situated outside outer level of ad1+ad2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ evenly sclerotized, forming distinct frame. Rudiment of process for attachment of dilators of anal valves (situated in region of anterior cuticular suckers) mediad to marginal sclerotized border of attachment organ. Lateral margins of attachment organ distinctly separated anteriorly. Alveolar sclerite of ps2 with weakly sclerotized edges, edges not straight. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple. Genual setae mG I shorter than genu I, mG II distinctly shorter than leg II, but longer than femur II. Tarsal setae la I-II distinctly longer than famulus. Tarsal setae ra I-II not bifid. Tarsal setae ra I-II spiniform. Tarsal setae wa I-II and s III nearly needle-like, slightly widened basally and with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical. Alveoli of tarsal setae d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterio-proximal lobe of ambulacral stalk distinctly elongated, with sides more then 2 times longer than base. Posterior condylophore present. Anterior condylophore I-II bent distally. Setae mG II widened and flattened. Setae vF II widened and flattened. Seta d III situated close to tarsal base, distance to base shorter than diameter of d III alveolus. Leg IV extending to posterior edge of hysterosoma. Tarsus IV not enlarged, about 3 times shorter than width of trochanter IV. Tarsus IV fused to tibia dorsally. Setae w IV thick, almost as thick as 1/2 of d IV width, distinctly longer than leg IV; situated at base of tarsus IV. Setae f IV short. Setae s IV absent. Setae wF IV distinctly extending beyond apex of tarsus IV, longer than leg IV. 804e96d6de1dba40eb4e97de2cf3f062 taxon_sennertia_lauta http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Species Sennertia lauta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription Gnathosomal solenidia longer than distance between them (shorter in all other species of the horrida-group). Setae f2 situated lateral to hysterosomal shield (on hysterosomal shield in S. horrida and S. madagascarensis, variabale in S. potanini). Tarsal setae ra I-II spiniform, massive (filiform in S. madagascarensis , blade-like in S. horrida and S. potanini). Setae vF II widened and flattened (nearly filiform in all other species of the horrida-group). The new species is closely related to Sennertia ratiocinator sp. nov. but differs by the thinner setae 1a and longer setae si, se, cp, and e2 (Table 3). None of the measurements alone can be used for complete discrimination between the two species. The sum of the ratios, width of 1a/width of se and length of idiosoma/length of si is 2.7-3.3 (3.0±0.15, n=54), whereas in S. ratiocinator, it is 3.8-4.9 (4.2±0.25, n=36). A canonical variates analysis based on these four, size-corrected (Darroch and Mosimann 1985), variables resulted in 100% classification accuracy (not reported here). f0d53a8ba5424d27e8947cb0689e5db9 taxon_sennertia_lauta http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia lauta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Philippines (type locality), Malaysia. a85c3dbcc2e6dc6f649ccb8ff4ef5917 taxon_sennertia_lauta http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia lauta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Zonohirsuta) fuliginata Pérez, 1901 (type host); Xylocopa (Zonohirsuta) dejeanii Lepeletier, 1841. 907a02fe138a4bbc87e60d7feba1d747 taxon_sennertia_lauta http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia lauta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology On host females, the mites attach to the groove between scutellum and metanotum, whereas on males they are situated on the anterior scutum (see Figs below). 5aa1b26dad999d9d28ca83ae7c0eac7a taxon_sennertia_leclercqi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia leclercqi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Sud-Kivu, Uvira (type locality) and Lake Albert 6383ba4775a850206d4b892059db8271 taxon_sennertia_leclercqi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia leclercqi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Xenoxylocopa) inconstans Smith, 1874 (as Mesotrichia inconstans) e42fb8a0e0084a07862661c13f5c5d8c taxon_sennertia_leei http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia leei, phoretic deutonymph, ex Xylocopa bombylans, Australia, BMOC 86-1022-016 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_leei_BMOC_86-1022-016.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e dfb0f60d0ff3700783df8acc1944dbcd taxon_sennertia_leei http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia leei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia: New South Wales (Kuring-gai) (as "Ku-rin-gai") (type locality) (Fain, 1982), Queensland, South Australia (Klimov & OConnor, 2008) f6823efaa7b6fdeb50e70786277fc05a taxon_sennertia_leei http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia leei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Lestis) bombylans (Fabricius, 1775) (as Lestis bombylans) (type host); Xylocopa (Lestis) aerata (Smith, 1851) (IT IS name Xylocopa aeratus (Smith, 1851)); (Fain, 1982; Klimov & OConnor, 2008) b9ad747a4a7d9afc9cdf0da40d68fe73 taxon_sennertia_longipilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia longipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina: Misiones Prov. 87e0e4a4adff9d21e748af44c7882a0f taxon_sennertia_longipilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia longipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Schonnherria) splendidula splendidula 8594266e17ba8b515d4a3345eda907d9 taxon_sennertia_loricata http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia loricata, phoretic deutonymph (holotype), ex Xylocopa micans, Florida, BMOC 04-0804-001 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_loricata_holotype_BMOC_04-0804-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e edeea23410e8f4cbb059870bbb7bf442 taxon_sennertia_loricata http://purl.org/dc/dcmitype/StillImage image/jpeg holotype slide http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_loricata_holotype_slide_BMOC_04-0804-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3a35135b74b63690dec1cce74d05e16d taxon_sennertia_loricata http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Sennertia loricata dispersing on the bee Xylocopa micans, Florida BMOC 03-1106-082 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_loricata_on_host_BMOC_03-1106-082.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 0c1bfc5559d7e7f9236507b6d684994f taxon_sennertia_loricata http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia loricata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield formed by long striae, with sclerotization, sclerotization situated between striae. Distinct rudiments of vi present. Setae si distinctly posterior se. Setae si exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 distinctly longer than d1-h1, nearly as long as se, situated on anterior edge of hysterosomal shield. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated outside hysterosomal shield. Setae 4b, g, and 4a without distinct rhomb-like widening filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2), anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers absent. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple, mG II almost as long as leg II or longer. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, blade-like. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical, d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore absent. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV distinctly protruding apex of tarsus IV.
Other instars unknown. 5bde46c8f39b569d637daa5a6fec23a7 taxon_sennertia_loricata http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia loricata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Florida, Louisiana, Puerto Rico, Texas, Virginia; Mexico: Chiapas, Jalisco. 84e560ad918f33c5e0c5fd8d664f4e21 taxon_sennertia_loricata http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia loricata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Schonnherria) micans, Xylocopa (Schonnherria) barbatella barbatella, Xylocopa (Schonnherria) muscaria. One specimen found on Xylocopa (Neoxylocopa) mordax. 379aed2c9523466f5d7d88a3bef77744 taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/StillImage image/jpeg Sennetia lucrosa, phoretic deutonymph (holotype), ex Xylocopa varipuncta, Califorornia, BMOC 03-0630-038 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennetia_lucrosa_holotype_BMOC_03-0630-038.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7ce731547b3e61b8425a89730b685680 taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia lucrosa dispersing on the carpenter bee, Xylocopa californica arizonensis, Arizona, BMOC 03-0904-004 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_lucrosa_on_host_BMOC_03-0904-004.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c240bf2e8ee9311fc9d4533101e5ca61 taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia lucrosa dispersing on the carpenter bee Xylocopa varipuncta, California, BMOC 03-0630-036 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_lucrosa_on_host_BMOC_03-0630-036b.JPG http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2ee96c6c630a86d77328dfc22b4ec54a taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia lucrosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se; exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 subequal to or less than 1. 3 times longer than d1; distinctly longer than h1; long, nearly as long as se; situated anterior to hysterosomal shield. Setae d1 and e1 distinctly longer than h1; d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2; not touching hysterosomal shield. Lateral gland openings situated outside hysterosomal shield. Setae 4b, g, and 4a without distinct rhomb-like widening, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers absent. Longitudinal hysterosomal sclerite present, short. Ventral hysterosoma smooth. Genual setae mG I-II simple. Genual setae mG II distinctly shorter than leg II, but longer than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II bifid, blade-like. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical, not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV distinctly not reaching base of tarsus IV. Ratio tarsus IV/anterior suckers 1.2-2.1 (1.6±0.19, n=72); ratio tarsus IV/hT II 0.7-1.3 (0.9±0.11, n=72).
Other instars unknown.
Notes. Similar to Sennertia faini Baker and Delfinado-Baker, 1983 (see key to species). See also notes below.
We constructed CVA models for S. lucrosa (n=72, all known localities and hosts) and Sennertia faini (n=6, type series). The models including 2-10 log-transformed Darroch and Mosimann shape variables separate the two with 100% accuracy in both analysis and jackknife cross-validation. Ratios created by using raw variables with the highest and the lowest correlation with the canonical function allow for almost complete bivariate discrimination (see key to species above). Sennertia faini is only known from the type series collected from the honey bee from Guatemala. We do not have any mites similar to either S. lucrosa or S. faini s. str. from Central America, so the relationships between the two morphospecies and possible host effect in S. faini cannot be determined. However, the distribution of Xylocopa guatemalensis and the occurrence of S. lucrosa on this host in central Mexico might indicate that the differences of S. faini and S. lucrosa are influenced by host effect of the former, and therefore both belong to the same species.
Correlation of heteromorphic deutonymphs and feeding instars is based on the sharing of the same host species and relative abundance of S. lucrosa deutonymphs. ce7429c321b2d9ce5dc890705cbb56fa taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia lucrosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, California (type locality), Texas; Mexico: Baja California, Baja California Sur, Colima, Chiapas, Morelos, Puebla, Sonora. 3b421a6b4ae80450ee5c9f98b24c2d92 taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia lucrosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) varipuncta (type host), Xylocopa (Neoxylocopa) clarionensis, Xylocopa (Neoxylocopa) mexicanorum, Xylocopa (Notoxylocopa) tabaniformis orpifex, Xylocopa (Notoxylocopa) guatemalensis, Xylocopa (Xylocopoides) californica arizonensis, Xylocopa (Xylocopoides) californica, Xylocopa (Xylocopoides) californica diamesa. b83c8693c27173983f70707b7ad8b9b8 taxon_sennertia_lucrosa http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia lucrosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology This mite species may destroy a small percentage of X. tabaniformis orpifex and X. varipuncta larvae in their nests (Nininger, 1916). Such behavioral features of the bee hosts as use of common surface entrance for several intraspecific or interspecific nest tunnels (X. t. orpifex and X. varipuncta) and consuming provision from adjoining nests by newly emerging bees (X. t. orpifex) (Cruden, 1966; Nininger, 1916) may facilitate mite dispersal. 3d6962a30ca1534b10aefbfcdd11a074 taxon_sennertia_madagascarensis http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia madagascarensis, phoretic deutonymph (holotype), ex Xylocopa sp., Madagascar, BMOC 74-0729-005 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_madagascarensis_BMOC_74-0729-005.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 8b852fd29ec539dc011ffc1fbc4f026d taxon_sennertia_madagascarensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia madagascarensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Madagascar: Antananarivo (Antananarivo) (as "Tananarive") (type locality) 97f80628b178e43d7bd109c1d6a9ef9b taxon_sennertia_madagascarensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia madagascarensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Prosopoxylocopa) mirabilis Hurd and Moure, 1963 (type host); Xylocopa (Koptortosoma) calens Lepeletier, 1841; (Fain, 1971; Klimov et al., 2007b) 78fe5bf40fee7ba7c604b7ddb585f97a taxon_sennertia http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite genus Sennertia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Notes. Although the monophyly of Sennertia is well supported (Klimov & OConnor, 2008), the internal relationships among different lineages and, especially their early divergence, are not clear. Sennertia zhelochovtsevi represents a mixture of advanced (the shape of hysterosomal shield) and plesiomorphic characters (relative length of si and c1), and the latter indicate its early divergence. Several characters (length of setae c1 and the reduction of hysterosomal sclerotization) suggest its possible sister-group relationship with the New World clades associated with Xylocopa.
With the extremely conservative and variable morphology in several distant lineages, the question about the monophyly of some Ceratina and Xylocopa associated lineages remains unsolved. The high rate of convergent morphological changes probably occurred because of correlated increase of the lengths of setae situated off the hysterosomal shield following its reduction. A preliminary phylogenetic analysis of several putative groups is given by Klimov and OConnor (2008).
Different groups of Sennertia phoretic deutonymphs are diagnosed in the following key (excludes the vaga-group, which is based on adults only):

Key to subgenera and species-groups of Sennertia
(phoretic deutonymphs, unique character states underlined)

1 Setae c1 long, nearly as long as se. Setae si nearly as long as se ... 2- Setae c1 shorter than se, microsetae. Setae si variable... 8 2(1) Hysterosomal shield triangle. Opisthosomal gland openings outside shield. Tarsal setae w IV short. Posterior apodeme IV absent. Xylocopa (Proxylocopa). Mediterranean, Middle Asia ... zhelochovtsevi-group (new)- Hysterosomal shield not triangle. Position of opisthosomal gland openings, length of tarsal setae w IV, and presence/absence of posterior apodeme IV variable. New World. (Amsennertia s. l.)... 3 3(2) Tarsal ventral setae w IV long, distinctly longer than leg IV. Posterior apodeme IV present. Setae 1a, 3a, 4b conoidal; c3, 4a, and g inflated at bases. Setae wa I-II bifid ... devincta-group - Tarsal ventral setae w IV distinctly shorter than leg IV. Posterior apodeme IV absent. Setae 1a, 3a, 4b, c3, 4a, g, and wa I-II simple. Associated with Xylocopa. (Amsennertia s. str.)... 4 4(3) Setae d1, e1, and h1 nearly uniform in length, microsetae, d1 and e1 distinctly shorter than respective 1/2 of distance between them. (-) Setae c1 situated on anterior margin of hysterosomal shield (frontalis-, loricata-groups) or anterior to it (americana-group). Opisthosomal gland openings on (frontalis-group) or outside hysterosomal shield (loricata-group, americana-group)... 6 - Setae d1 and e1 distinctly longer than h1, at least one of them longer than 1/2 of distance between bases. Setae c1 anterior to hysterosomal shield. Opisthosomal gland openings outside hysterosomal shield... 5 5(4) Setae mG II almost as long as leg II, distinctly longer than vF II. Tarsal setae ra I-II simple ... ignota-group- Setae mG II distinctly shorter than leg II and setae vF II. Tarsal setae ra I-II bifid ... faini-group6(4) Setae c1 anterior to hysterosomal shield ... americana-group- Setae c1 situated on anterior margin of hysterosomal shield... 7 7(6) Opisthosomal gland openings outside hysterosomal shield. Usual striate pattern of hysterosoma accompanied by sclerotization ... loricata-group- Opisthosomal gland openings on hysterosomal shield. Striate pattern of hysterosoma without sclerotization ... frontalis-group8(1) Caruncle I-III with 2 distinct transparent lobes. Setae si distinctly longer than se. Tarsal setae e and f I-II strongly asymmetrical, f about 2 times shorter and filiform, e longer and lanceolate. (-) Hysterosomal shield triangle in outline. Setae pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II spiniform. Associated with Xylocopa in the Neotropics ... Spinosennertia Fain- Caruncle I-III with 1, usually sclerotized lobe (distal lobe absent). Tarsal setae e and f I-II slightly asymmetrical, f maximum 1.5 shorter than e. Other characters variable... 9 9(8) Setae c1 anterior to hysterosomal shield. Associated with Xylocopa. Afrotropical, Oriental, Eastern Palaearctic ... Afrosennertia Fain (=Asiosennertia Fain)- Setae c1 situated on hysterosomal shield. Associated with Xylocopa and Ceratina. Old World (Sennertia)... 10 10(9) Setae si of medium length, not microsetae... 11 - Setae si microsetae... 12 11(10) Ventral tarsal seta w IV long, distinctly longer than tarsus IV. Setae si and se on same transverse level (si distinctly posterior in one undescribed species). Associated with Xylocopa. Oriental region and Madagascar ... horrida-group- Ventral tarsal seta w IV short, usually shorter than tarsus IV. Setae si posterior to se. Associated with Ceratina. Neotropics ... surinamensis-group12(10) Ventral tarsal seta w IV elongated. Afrotropic and Oriental, Eastern Palaearctic. Associated with Xylocopa (placement of the African clade here is questionable) ... japonica-group- Ventral tarsal seta wa IV not elongated. Associated with Xylocopa or Ceratina. Old World. (probably paraphyletic) ... cerambycina-group (including Eosennertia Kurosa) 766c5c99d6918e9d0d69af5de51e0aa9 taxon_sennertia_mesotrichia http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia mesotrichia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Bambesa (type locality) and other localities, including Katompi and Ituri 148e464eeb55e3fdcd10125b5f7edeb7 taxon_sennertia_mesotrichia http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia mesotrichia http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) africana (Fabricius, 1781) (type host) (as Mesotrichia africana) (Bambesa, "several other localities"); Xylocopa (Koptortosoma) lepeletieri Enderlein, 1903 (as Mesotrichia lepeletieri) (Katompi); Xylocopa (Xylomelissa) hottentotta Smith, 1854 (as Xylocopa tarsata) (Ituri); (Fain, 1971; Fain, 1981a) 4c6d35af88614a27b7f1f1d607fbb0a6 taxon_sennertia_micheli http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia micheli http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java 5aaf39d4a1aba58f553821a63cc1937c taxon_sennertia_micheli http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia micheli http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Bembix borrei Handlirsch, 1893 (Hymenoptera: Sphecidae) 445e4eccc3a47b64325de9199471ffb3 taxon_sennertia_monicae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia monicae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Sud-Kivu (Uvira) (type locality) cca35d16b17f24b1cfef1bb238533b26 taxon_sennertia_monicae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia monicae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Xenoxylocopa) inconstans Smith, 1874 (as Mesotrichia inconstans) 90d377108bad6bcbceb468daba399c9b taxon_sennertia_morstatti http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia morstatti, phoretic deutonymph, ex Xylocopa nigrita, Cameroon, BMOC 90-1212-018 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_morstatti_BMOC_90-1212-018.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 57266038cb12c1ac9b18f570072ae0dd taxon_sennertia_morstatti http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia morstatti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tanzania: Amani ("Amani, Deutsch-Ostafrica") (type locality) (Vitzthum, 1914); Democratic Republic of the Congo; Sudan; (Fain, 1981a); Cameroon (our data). d62e0300f37023221c65c1166b597edf taxon_sennertia_morstatti http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia morstatti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) nigrita (Fabricius, 1775) (as Koptorthosoma nigrita) (type host); Xylocopa (Koptortosoma) sp. (as Xylocopa (Koptorthosoma) sp.) (Sudan); Megachilidae (Congo); (Vitzthum, 1914; Fain, 1981a) 7d8a136cf5a58e406d3150056e1d16a4 taxon_sennertia_oudemansi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia oudemansi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Japan (no specific location) dd8b0dbc7beecc131d7ef0f40a54a6b0 taxon_sennertia_oudemansi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia oudemansi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Biluna) nasalis Westwood, 1842 (as Xylocopa dissimilis). 666eb4bb406257c8731683fd3bf5c0c4 taxon_sennertia_perturbans http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia perturbans http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tanzania: Amani ("Amani, Ostafrica") 9c197dae8d6744abb70bd9dd582e31a4 taxon_sennertia_perturbans http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia perturbans http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) nigrita (Fabricius, 1775) (as Koptorthosoma nigrita) 42656f925e29a1754f5746d5a5798ccd taxon_sennertia_pirata http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia pirata, phoretic deutonymph (holotype), ex Xylocopa sp., Bahamas, BMOC 04-0508-310 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_pirata_holotype_BMOC_04-0508-310.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b20ce8780708cfee8e4d65ca19434d55 taxon_sennertia_pirata http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia pirata dispersing on the propodeum of the bee, Xylocopa mordax, US Virgin Islands http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_pirata_on_host_BMOC_04-1122-011a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9d031c5dcb41cb3992ef7f9ca285dbcc taxon_sennertia_pirata http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia pirata dispersing on the propodeum of the bee, Xylocopa mordax, US Virgin Islands http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_pirata_on_host_BMOC_04-1122-011c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 1801432c3f0f7f81cbcdb8d9cea2d825 taxon_sennertia_pirata http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia pirata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 distinctly longer than d1-h1, long, nearly as long as se, situated on anterior edge of hysterosomal shield. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 present. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated on hysterosomal shield. Setae 4b, g, and 4a with rhomb-like widening (sometime weakly-developed). Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, and ra I-II filiform or nearly filiform (wa I-II spiniform). Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2), anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma distinctly striated. Genual setae mG I-II simple, mG II almost as long as leg II or longer. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, blade-like. Tarsal setae wa I-II and s III spiniform (at least s III with rounded apices). Tarsal setae d I-II distinctly widened, lanceolate. Tarsal setae d and f I-II almost symmetrical, not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV not reaching apex of tarsus IV. 9a2e451a4dae7249fd89327d718f6040 taxon_sennertia_pirata http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia pirata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Bahamas, British Virgin Islands, Cuba, USA: Puerto Rico, Virgin Islands of the United States. 2dc865a51577dbbd20c3a51963b0f4c7 taxon_sennertia_pirata http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia pirata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Neoxylocopa) cubaecola, Xylocopa (Neoxylocopa) mordax, Xylocopa sp. e23671b10ea369b58bc6ae8e6423b0f4 taxon_sennertia_potanini http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia potanini, phoretic deutonymph (lectotype), ex Xylocopa tranquebarorum (or X. rufipes), China, BMOC 11-0618-004 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_potanini_lectotype_dors_BMOC_11-0618-004.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 71ad7e9f668b0a5cd9d36fa179d6406a taxon_sennertia_potanini http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia potanini, lectotype heteromorphic deutonymph, ventral http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_potanini_lectotype_ventr_BMOC_11-0618-004.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7c13ec44e26ca4bc750c402581dc5044 taxon_sennertia_potanini http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia potanini, lectotype slide http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_potanini_lectotype_slide_BMOC_11-0618-004.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 467936f138a3fdf57c2c7a7e0dd172a9 taxon_sennertia_potanini http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia potanini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution China: Sichuan (Yazhou) 3081cae2669bbc3c6b8c64e0a61875da taxon_sennertia_potanini http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia potanini http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Biluna) tranquebarorum (Swederus, 1787) or Xylocopa (Bomboixylocopa) rufipes Smith, 1852 (as Xylocopa pictifrons) 693d4e7f708d900c066a32e71171397a taxon_sennertia_queenslandica http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia queenslandica, phoretic deutonymph (holotype), ex Xylocopa bryorum, Australia, BMOC 05-0420-081 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_queenslandica_BMOC_05-0420-081.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e b8529aab78b412bc5cc819c6833a3ddb taxon_sennertia_queenslandica http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia queenslandica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Australia: Queensland (Moa Island) ("Moa Id., Torres Straits") c0666d0b35f236df0532f834744cb104 taxon_sennertia_queenslandica http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia queenslandica http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) bryorum (Fabricius, 1775) (as Mesotricha bryorum) 2cc22086174ef728cce6c8de151ff330 taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia ratiocinator, phoretic deutonymph (holotype), ex Xylocopa bhowara, Sri Lanka, BMOC 05-0420-066 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_ratiocinator_holotype_BMOC_05-0420-066.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 71ccbe8c059980ac3cea18409aed69d2 taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia ratiocinator dispersing in the groove between scutellum and metanotum of the female carpenter bee, Xylocopa bhowara, Sri Lanka, BMOC 05-0420-066 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_ratiocinator_on_host_BMOC_05-0420-066.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bee5fd93f5bfc76a0f49f0317d3fa2cf taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia ratiocinator dispersing in the groove between scutellum and metanotum of the female carpenter bee, Xylocopa bhowara, Sri Lanka, BMOC 05-0420-066 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_ratiocinator_on_host_BMOC_05-0420-066d.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e bb1116473675fa5de7e01f25e78e1de9 taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/Text text/html Diagnostic Description of the Bee Mite Species Sennertia ratiocinator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription The new species is closely related to Sennertia lauta but differs by thicker setae 1a and shorter setae si, se, cp, and e2 (Table 3). None of the measurements alone can be used for a complete discrimination between the two species. The sum of ratios width of 1a/width of se and length of idiosoma/length of si is it is 3.8-4.9 (4.2±0.25, n=36), whereas in S. lauta it is 2.7-3.3 (3.0±0.15, n=54). A canonical variates analysis based on these four, size-corrected, variables resulted in 100% classification accuracy (not reported here). 4963bd50964fa40518d7791b92fd0e9c taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia ratiocinator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Sri Lanka. 2af358884a7115fd6bfcd196ebdddb99 taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia ratiocinator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Zonohirsuta) bhowara Maa, 1938 (type host); Xylocopa (Nodula) amethystina (Fabricius, 1793); Xylocopa (Zonohirsuta) dejeanii Lepeletier, 1841. 692f523e7a71f257f5d1fc48ac0b3efd taxon_sennertia_ratiocinator http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia ratiocinator http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology On female hosts, mites attach to the groove between scutellum and metanotum, whereas on males they are situated on the anterior scutum. 510dfea165c46449b30c90060c147b0b taxon_sennertia_recondita http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia recondita, phoretic deutonymph (holotype), ex Ceratina eximia, Mexico, BMOC 96-0510-231 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_recondita_holotype_BMOC_96-0510-231.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e d45ed9cfd77b6b4139a0328d2b9d484a taxon_sennertia_recondita http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia recondita dispersing on the propodeum of the small carpenter bee, Ceratina eximia, Mexico, BMOC-96-0510-231 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_recondita_on_host_BMOC-96-0510-231b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 25ae5e5a6c489fb0a9f399164830f530 taxon_sennertia_recondita http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia recondita http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width, gnathosomal solenidia exceeding half of setae vi. Supracoxal setae scx situated on dorsal extension of posterior apodeme I. Hysterosomal shield distinctly expanded beyond lateral gland opening and bases of setae f2. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield formed by long striae, with sclerotization, sclerotization shifted to posterior striae. 10-14 lines between setae se and si. Distinct rudiments of vi absent. Setae si distinctly posterior se; exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 and d1-h1 uniform in length, microsetae. Setae c1 microsetae; situated posterior to anterior margin of hysterosomal shield. Setae c3 nearly spiniform, situated on soft cuticle. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated on hysterosomal shield. Setae 4b filiform. Setae 4b, g, and 4a without distinct rhomb-like widening, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV present, connected to anterior apodeme III. Additional posterior sclerite of posterior apodeme IV present. Conoids ps2 anterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II bifid at tips, mG II shorter than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, filiform. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II foliate. Tarsal setae d and f I-II almost symmetrical, d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV not protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV not protruding tibia IV.
Other instars unknown.
Notes. Similar to S. surinamensis and S. sodalis. See key above for the diagnostic characters. bcf13b7a75885e7d3be2713235dd3fdb taxon_sennertia_recondita http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia recondita http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Quintana Roo (type locality), Belize: Cayo. 8bff8ec0bf38f6a062b5a14ad95ffeff taxon_sennertia_recondita http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia recondita http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Calloceratina) eximia. 6c577fff1bc478026caedcd404133f32 taxon_sennertia_robusta http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia robusta, phoretic deutonymph, ex Xylocopa fenestrata, India, BMOC 05-0102-040 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_robusta_BMOC_05-0102-040.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c19e0cbb4722424d15b1e3153624c0ab taxon_sennertia_robusta http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia robusta dispersing on the the wings of the bee Xylocopa fenestrata, India, BMOC 05-0102-040 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_robusta_on_host_BMOC-05-0102-040.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 164fdc19e3fa9dc7693b8f5f7ecf2bc3 taxon_sennertia_robusta http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia robusta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution India, Haryāna (Hisār) ("Hissar") (type locality) (Delfinado & Baker, 1976), Tamil Nādu (Ramaraju & Mohanasundaram, 2001). 63ad5abd6c0ee68c31272d68ef898cd8 taxon_sennertia_robusta http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia robusta http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations "megachilid bee (PL-480; India: Hissar; probably in BLCU) (type host); Xylocopa sp. ("India", probably in USNM); Xylocopa (Nodula) amethystina (Fabricius, 1793) ; (Delfinado & Baker, 1976; Ramaraju & Mohanasundaram, 2001) 1b9090a8fe37b8fe965c69714c2f96f5 taxon_sennertia_roepkei http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia roepkei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java (Bogor) (as "Buitenzorg") 631e3e62578fa443de560e63205c2632 taxon_sennertia_roepkei http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia roepkei http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations On eggs of Horia (as "Cissites (= Horia)") (Coleoptera: Meloidae) in nest of Xylocopa sp. 590bbcb1dfd0be133eba3b4da4bc0f8c taxon_sennertia_sayutara http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia sayutara, phoretic deutonymph (holotype), ex Ceratina laticeps, Costa Rica, BMOC 04-0508-284 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sayutara_holotype_BMOC_04-0508-284.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e da33a0ce89d1f233cecd7d87577c7d86 taxon_sennertia_sayutara http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia sayutara around the metasomal acarinarium of the bee Cereatina laticeps, Costa Rica, BMOC 04-0508-284 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sayutara_on_host_BMOC-04-0508-284a.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a948e72d54f9bcef44112f0df6f3c678 taxon_sennertia_sayutara http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia sayutara around the metasomal acarinarium of the bee Cereatina laticeps, Costa Rica, BMOC 04-0508-284 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sayutara_BMOC_04-0508-284.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a3566848b9a258536b8f7b14f183a700 taxon_sennertia_sayutara http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia sayutara http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia distinctly shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield distinctly expanded beyond lateral gland opening and bases of setae f2. Lateral edges of hysterosomal shield in anterior part not narrowing, rounded. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeding diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization; at posterior end of hysterosomal shield nearly uniformly parallel. Striae long. Distinct rudiments of vi present, situated at level of se. Setae si distinctly posterior se. Setae si exceeding slightly longer than se, as thick as se. Setae c1 long, nearly as long as se, distinctly longer than d1-h1, situated on anterior edge of hysterosomal shield. Setae c3 nearly spiniform, situated on weakly developed, small sclerite. Setae cp not reaching im and setae e2. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2. Setae e2 not touching hysterosomal shield. Setae h3 non-applicable. Setae h3 non-applicable. Lateral gland openings situated on hysterosomal shield. Setae pR I-II, sR III, wF IV, gT I-II, kT III filiform. Setae 1a, 4b, and 3a conoidal; g and 4a spiniform, with attenuated tips. Posterior apodemes II and anterior apodemes III partially fused. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV present, not connected to anterior apodeme III. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2). Conoids ps2 anterior to ps1, situated outside outer level of ad1+ ad2. Suckers ad3 not enlarged, smaller than central suckers, without rough sclerotization. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present, very small. Posterio-medial longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple, shorter than leg II, but longer than femur II. Tarsal setae la I-II microsetae. Tarsal setae ra I-II not bifid, spiniform. Tarsal setae wa I-II spiniform, bifid apically, s III spiniform, simple. Tarsal setae d I-II foliate, shorter than f I-II. Tarsal setae d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Setae sR III not protruding femur III. Seta d III situated shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter than width of trochanter IV. Setae w IV almost as 1/2 of d IV width, distinctly longer than leg IV. Setae w IV situated on middle of tarsus IV. Setae s IV present, very short. Setae wF IV not protruding tibia IV. Other instars unknown. d1f49ab2e4a88d846667efe906762572 taxon_sennertia_sayutara http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia sayutara http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Costa Rica: San Jose. 0660570a6090201bfa53e9dd6bb847ad taxon_sennertia_sayutara http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia sayutara http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina laticeps. cfb3d1121481992000213cdfd7fd4cfc taxon_sennertia_sayutara http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Sennertia sayutara http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found in the metasomal acarinarium of the host. 46cf061bfcb9e46c61bd5c974d59b255 taxon_sennertia_scutata http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia scutata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution South Africa: former Cape Prov. (type locality) and Western Cape: Cape Town 045106929c7c09f957698930b09fa9a6 taxon_sennertia_scutata http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia scutata http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Dalyatina) truncata Friese, 1905 ("Cape") (type host); Ceratina (Pithitis) nasalis Friese, 1905 (as Ceratina viridior) (Cape Town) 69f803dd4c786727b63da7dba67c2a7b taxon_sennertia_segnis http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia segnis, phoretic deutonymph (holotype), ex Xylocopa californica californica, California, BMOC 03-0604-039 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_segnis_holotype_BMOC_03-0604-039.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 56297c0dea3a3f2e98f597c8cf435f3b taxon_sennertia_segnis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of Sennertia segnis on the carpenter bee, Xylocopa californica, Arizona http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_segnis_on_host_BMOC_04-1222-155.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 77f09dbe6ddf2aa15965428a0266b211 taxon_sennertia_segnis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia segnis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 subequal to or less than 1. 3 times longer than d1; distinctly longer than h1; long, nearly as long as se; situated anterior to hysterosomal shield. Setae d1 and e1 distinctly longer than h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated outside hysterosomal shield. Setae 4b, g, and 4a without distinct rhomb-like, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Posterior processes of coxal apodemes IV non-applicable. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers absent. Longitudinal hysterosomal sclerite present, short. Ventral hysterosoma smooth. Genual setae mG I-II simple, mG II distinctly shorter than leg II, but longer than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II bifid, blade-like. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II slightly widened. Tarsal setae d and f I-II almost symmetrical, not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV almost reaching or slightly protruding apex of tarsus IV.
Other instars unknown (tritonymph was observed inside pharate HDN but it cannot be fully described). 5473d637809845485ae2cba6dec1fdd2 taxon_sennertia_segnis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia segnis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, California (type locality); Mexico: Michoacán de Ocampo, Puebla. 8a19c3841486c912be3d9ac718e6dfd9 taxon_sennertia_segnis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia segnis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Xylocopoides) californica, Xylocopa (Xylocopoides) californica arizonensis, Xylocopa (Xylocopoides) californica diamesa, Xylocopa (Xylocopoides) cyanea, Xylocopa (Notoxylocopa) tabaniformis orpifex. d743f51baf8c87b77b6a2472c4068d3d taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia shimanukii, phoretic deutonymph (holotype), ex Xylocopa mexicanorum, Texas, BMOC 05-0420-075 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_shimanukii_BMOC_05-0420-075.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 3d66fb39ed406109eedd8bc13855374b taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia shimanukii dispersing on the bee, Xylocopa sonorina, California http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_shimanukii_on_host_BMOC_05-0420-365.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e caa8c52afc1a07bdfc3abc44050a3030 taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia shimanukii dispersing on the bee, Xylocopa sonorina, California http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_shimanukii_on_host_BMOC_05-0420-365c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2439041f62b8450243ac89070a00e16b taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia shimanukii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width. Supracoxal setae scx situated on separate small sclerite. Hysterosomal shield lateral gland openings and bases of f2 nearly on edge of hysterosomal shield, or the former outside the shield. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield without sclerotization, formed by long striae. Distinct rudiments of vi present. Setae si distinctly posterior se, exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 distinctly longer than d1-h1; long, nearly as long as se; situated on anterior edge of hysterosomal shield. Setae d1 and e1 nearly uniform in length with h1; situated on hysterosomal shield. Sclerite between ia and d2 present. Setae e2 subequal with d2; not touching hysterosomal shield. Lateral gland openings situated on hysterosomal shield. Setae 4b, g, and 4a with rhomb-like widening (sometime weakly-developed). Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II filiform. wa I-II spiniform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted; almost straight. Posterior apodeme IV absent. Conoids ps2 posterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II simple, mG II distinctly shorter than leg II, but longer than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, filiform. Tarsal setae wa I-II and s III spiniform (at least s III with rounded apices). Tarsal setae d I-II distinctly widened, lanceolate. Tarsal setae d and f I-II almost symmetrical, d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated close to tarsal base, distance usually subequal or shorter than diameter of d III alveolus. Leg IV protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV slightly protruding apex of tarsus IV.
Other instars unknown.
Notes. Similar to Sennertia augustii and S. frontalis (see p. 84) 00048afc7dfb2f454474003bc16fe47e taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia shimanukii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Guatemala: Sololá (type locality); Mexico: Baja California Sur, Nayarit, Veracruz; USA: California, Texas. 7544935d60303e5e68b2a1eaaf6d663b taxon_sennertia_shimanukii http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia shimanukii http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Apis mellifera (type host, accidental); Xylocopa (Neoxylocopa) varipuncta, Xylocopa (Neoxylocopa) mexicanorum, Xylocopa (Neoxylocopa) sonorina, Xylocopa (Xylocopoides) californica arizonensis, Xylocopa (Neoxylocopa) fimbriata (Apidae). 795717814c2c1449673b13faaeda6223 taxon_sennertia_simplex http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia simplex, phoretic deutonymph, ex Xylocopa nigrita, Africa, HK 85-0115-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_simplex_HK_85-0115-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 60cc6367b8818cd4f8f23fbd883d145a taxon_sennertia_simplex http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia simplex http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Cameroon 32e34483e5a898fe3f69c74864d8a39f taxon_sennertia_simplex http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia simplex http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Koptortosoma) nigrita (Fabricius, 1775) 6b94bab006be42fb96ce49cef91ba71a taxon_sennertia_sodalis http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia sodalis, phoretic deutonymph (holotype), ex Ceratina sp., Mexico, BMOC 03-1008-055 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sodalis_holotype_BMOC_03-1008-055.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 798570b02d910a9ca56940cec85c27e0 taxon_sennertia_sodalis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia sodalis dispersing on the propodeum of the samll carpenter bee, Ceratina eximia, Cost Rica, BMOC-96-0510-229 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sodalis_on_host_BMOC-96-0510-229c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c366b226c0f02bbc086ac79d44dc423f taxon_sennertia_sodalis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia sodalis dispersing on the propodeum of the samll carpenter bee, Ceratina eximia, Cost Rica, BMOC-96-0510-229 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_sodalis_on_host_BMOC-96-0510-229.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e dc3a36d397830799016a99ccb70cb28b taxon_sennertia_sodalis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia sodalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic deutonymph. Gnathosomal solenidia shorter than 1/3 of femur I width, distinctly shorter than half of setae vi. Supracoxal setae scx situated on dorsal extension of posterior apodeme I. Hysterosomal shield distinctly expanded beyond lateral gland opening and bases of setae f2. Lateral edges of hysterosomal shield in anterior part not narrowing. Dorsal hysterosomal pouch absent. Distance between anterior margin of hysterosomal shield and setae si exceeds diameter of si bases. Striate pattern of idiosomal cuticle outside hysterosomal shield formed by long striae, with sclerotization, sclerotization shifted to posterior striae. 5-7 lines between setae se and si.. Distinct rudiments of vi absent. Setae si distinctly posterior se; exceed 1/2 of se, almost as thick as se. Diameter of si exceeds 1/2 of diameter of se. Setae c1 and d1-h1 uniform in length, microsetae. Setae c1 microsetae; situated posterior to anterior margin of hysterosomal shield. Setae c3 nearly spiniform, situated on soft cuticle. Setae d1 and e1 nearly uniform in length with h1. Setae d1 situated on hysterosomal shield. Sclerite between ia and d2 absent. Setae e2 subequal with d2, not touching hysterosomal shield. Lateral gland openings situated on hysterosomal shield. Setae 4b filiform. Setae 4b, g, and 4a without distinct rhomb-like widening, filiform. Setae 4b, pR I-II, sR III, wF IV, gT I-II, hT I-II, kT III, ra I-II, and wa I-II filiform. Posterior apodemes II and anterior apodemes III free. Anterior apodemes IV not interrupted, almost straight. Posterior apodeme IV present, connected to anterior apodeme III. Additional posterior sclerite of posterior apodeme IV absent. Conoids ps2 anterior to anterior transverse level of central suckers (ad1+2); anterior to ps1, situated outside outer level of ad1+2. Transparent margin of anterior suckers (ad3) without rough sclerotization. Suckers ad3 not enlarged, smaller than central suckers. Posterior and lateral borders of attachment organ not forming distinct frame. Sclerotized rudiment of anterior cuticular suckers present. Longitudinal hysterosomal sclerite present, long. Ventral hysterosoma smooth. Genual setae mG I-II bifid at tips, mG II shorter than femur II. Tarsal setae la I-II longer than famulus ε. Tarsal setae ra I-II not bifid, filiform. Tarsal setae wa I-II and s III filiform, needle-like, or widened basally but with attenuated end. Tarsal setae d I-II foliate. Tarsal setae d and f I-II almost symmetrical, d and f I not touching. Solenidion ω3 closer to f I than to ω1. Posterior condylophore present. Anterior condylophore I-II with distal bending. Seta d III situated shifted from tarsal base, distance distinctly exceeding diameter of d III alveolus. Leg IV not protruding posterior edge of hysterosoma. Tarsus IV not enlarged, shorter or less than 2 times longer than width of trochanter IV. Setae w IV thinner than d IV and distinctly shorter than leg IV, situated on middle of tarsus IV. Setae s IV present. Setae wF IV not protruding genu IV.
Other instars unknown. e49c7568dd938c428e59dd357283a2c0 taxon_sennertia_sodalis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia sodalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Mexico: Oaxaca, San Luis Potosí, Veracruz (type locality), Yucatán; Costa Rica, Panama. d7f65921e8d29c912af79ade1d6c7b1d taxon_sennertia_sodalis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia sodalis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina sp. , Ceratina (Calloceratina) eximia (Apidae). b3d32a11ddd6ef9eaea71ac0d5bceb9b taxon_sennertia_spinifera http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia spinifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo ("N.E. Lusambo, Zaïre") 213c0d85776c401c26ee2e93ad6448d6 taxon_sennertia_spinifera http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia spinifera http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Propithitis) aereola Vachal, 1903 af8884f0146a97af0abec8f7f4c1940b taxon_sennertia_splendidulae http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia splendidulae, phoretic deutonymph, ex Xylocopa splendidula, Argentina, HK 85-0107-005 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_splendidulae_HK_85-0107-005.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e da82d2fcf17499fb11a68110c37fa06f taxon_sennertia_splendidulae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia splendidulae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Argentina: Buenos Aires Prov. (La Plata) (type locality), Catamarca, Corrientes, Entre Ríos, Jujuy, La Pampa, Mendoza, Misiones, Río Negro, San Luis d6a75dd2809089dfc510e1aabac462af taxon_sennertia_splendidulae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia splendidulae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Schonnherria) splendidula Lepeletier, 1841 (type host); Xylocopa (Neoxylocopa) mendozana Enderlein, 1913 d5b79db2638b6790fc349b8926e78d3e taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia surinamensis, phoretic deutonymph, ex Ceratina chloris, French Guiana, BMOC 96-0510-228 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_surinamensis_BMOC_96-0510-228.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 05b83743126ba3d56626aa32cfd89c56 taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia surinamensis dispersing on the bee Ceratina chloris, French Guiana http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_surinamensis_on_host_BMOC-96-0510-228.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 81b8b14c85b97f4864fd1c2c7264815b taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia surinamensis dispersing on the bee Ceratina chloris, French Guiana http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_surinamensis_on_host_BMOC-96-0510-228c.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 7864f4d5a4cb3611856f2db8d6c963a9 taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia surinamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Known only from phoretic deutonymphs, see key. 42c4875cda42a0b73efa5703932221d2 taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia surinamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Suriname: Paramaribo (type locality); French Guiana, Panama. c8b5949e69f1f9320131e3732d31c7b7 taxon_sennertia_surinamensis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia surinamensis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Calloceratina) chloris. 45869747a5e46773ceceaf69396438a6 taxon_sennertia_tanythrix http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia tanythrix, phoretic deutonymph, ex Xylocopa torrida, Cameroon, BMOC 90-1212-014 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_tanythrix_BMOC_90-1212-014.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 13e2fff7c821de4b253b5aecfa6eed86 taxon_sennertia_tanythrix http://purl.org/dc/dcmitype/StillImage image/jpeg Phoretic deutonymphs of the mite Sennertia tanythrix dispersing on the the head of the bee Xylocopa torrida, Angola, BMOC 03-0630-005 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_tanythrix_on_host_BMOC-03-0630-005a http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c775e17ebf2b8d71ca13a9b3e47c2daf taxon_sennertia_tanythrix http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia tanythrix http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Haut-Congo Prov. "Uélé" (type locality) (Fain, 1971, 1981a); Angola; Cameroon (our data, unpubl.) 1868be74548894709c24f3299714823d taxon_sennertia_tanythrix http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia tanythrix http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) torrida (Westwood, 1838) (as Mesotrichia torrida) (type host); Xylocopa (Koptortosoma) imitator Smith, 1854 11b68a91e7eea8624660237a3bd59c3e taxon_sennertia_tunisiana http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia tunisiana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Tunisia: Silyānah (Maktar) (type locality), Nābul (Grombalia), Jundūbah (‘Ayn ad Darāhim, Tabarka), "Maletar", "Ain Sebaa" (probably in Morocco) 79ea0dcf96ebb4cd58a69fcf05083ea8 taxon_sennertia_tunisiana http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia tunisiana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Ceratina (Ceratina) cucurbitina (Rossi, 1792) (type host) (Maktar, Grombalia, "Ain Sebaa") ; Ceratina (Euceratina) albosticta Cockerell, 1931 ("Ain Sebaa"); Ceratina (Euceratina) mocsaryi Friese, 1896 (‘Ayn ad Darāhim); Ceratina dallatorreana Friese, 1896 (Tabarka); Ceratina (Euceratina) chalybea Chevrier, 1872 ("Maletar") 184f16a828cff2d8dea19db038c9f4f4 taxon_sennertia_vaga http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia vaga, female (holotype), ex Xylocopa tabaniformis orpifex, California, BMOC 04-1122-025 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_vaga_holotype_BMOC_04-1122-025.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e c8b6cc7ebb4d9028481ec4aebe561eca taxon_sennertia_vaga http://purl.org/dc/dcmitype/StillImage image/jpeg Females of the mite Sennertia vaga dispersing on the pronotum of the carpenter bee, Xylocopa tabaniformis orpifex, California (BMOC 05-0420-389) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_vaga_on_host_BMOC_05-0420-389b.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e dae7005ca2a970b212ea369ba344ae9a taxon_sennertia_vaga http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Species Sennertia vaga http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Phoretic and immobile deutonymph unknown, probably absent (see notes).
Adult. Supracoxal seta scx situated outside supracoxal sclerite, lateral to outer ridge of supracoxal sclerite. Alveoli of setae ve present. Dorsal idiosomal cuticle uniformly striate (striae may be with small tubercles). Dorsal cuticular pattern more or less uniform. Dorsal idiosomal setae c1-h1 filiform and long, reaching at least half of distance to next posterior pair of setae). Dorsal idiosomal setae cp, c3, h3 narrowing distally, not compressed dorso-ventrally, usually evenly barbed. Dorsal setae e2 and f2 filiform, nearly as long as h3. Prodorsal shield subquadrate, length/width 0.8-1.4. Prodorsal shield without falsifoveate pattern. Coxal fields III closed. Proximal acetabular extensions of ap' I partially border antiaxial margins of coxal fields I. Proximal acetabular extensions of ap' II partially border antiaxial margins of coxal fields II. Distal acetabular extensions of ap' II and ap'' II separate. Proximal acetabular extensions of ap' III partially border antiaxial margins of coxal fields III. Distal acetabular extensions of ap' III and ap'' III separate or not developed. Proximal acetabular extensions of ap' IV partially border antiaxial margins of coxal fields IV. Distal acetabular extensions of ap' IV and ap'' IV separate or not developed. Opisthosomal gland openings distinctly anterior to setae e2. Tarsal setae ra and la II absent. Solenidion ω2 I intermediate between subapical and submedial. Famulus ε lanceolate. Setae ba I as long as famulus ε or shorter. Setae ba II as long as famulus ε or shorter.
Female. Setae ad1 and ad2 present. Setae ps3 short, distinctly shorter than ps2; posterior to 4a level. External copulatory tube present. Setae h3 anterior to h2. Posterio-medial part of dorsal opisthosoma without distinct longitudinal linear pattern.
Male. Setae ad1 present. Genital setae spiniform; slightly (about on their diameter at base) anterior to progenital fold. Pseudanal setae ps3 outside progenital sclerites, filiform. Dorsal supporting sclerites short, as long as 2 diameters of aedeagus at base or shorter. Setae q I and p II present. Pretarsal suckers IV same as pretarsal suckers I-III.
Protonymph. Tarsal setae e IV absent; f IV present.
Larva. Proportional length of dorsal idiosomal setae as in other instars.
Notes. This species probably does not form phoretic deutonymphs. In the southern Nearctic, it is associated with hosts that also harbor S. lucrosa and S. shimanukii and sometimes may be found with the former on the same bee specimen. A single pharate tritonymph of S. lucrosa (BMOC 05-0420-076) is substantially distinct from tritonymphs of S. vaga in having la and ra II developed and coxal fields III opened. Feeding instars of S. augustii (deutonymphs virtually undistinguishable from S. shimanukii) also have opened coxal fields III. The same situation was also detected for some Neotropic Centris that have phoretic feeding instars of a similar species and heteromorphic deutonymphs of Centriacarus turbator. 6236a444035ea7b1b2767f97f50d534c taxon_sennertia_vaga http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia vaga http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA: Arizona, California; Mexico: Baja California Sur, Sinaloa. 1bb773108270ec70540feefaf4a90f0c taxon_sennertia_vaga http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia vaga http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Notoxylocopa) tabaniformis orpifex, Xylocopa (Notoxylocopa) tabaniformis androleuca, Xylocopa (Xylocopoides) californica. f440c951eb5126fee5af350e0c04eab9 taxon_sennertia_vanderhammeni http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia vanderhammeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Sri Lanka: Central (Peradeniya) ("Peradeniya. Ceylon") (type locality) 5e6a346d8ce2dfe8027625d6099aedb3 taxon_sennertia_vanderhammeni http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia vanderhammeni http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) tenuiscapa Westwood, 1840 (as Koptorthosoma tenuiscapa) 91a977c06f0b76496db9b65ef33d47df taxon_sennertia_vanderhammeni_brevipilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia vanderhammeni brevipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Java f6984efbdb22cd20377a1873a7df2425 taxon_sennertia_vanderhammeni_brevipilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia vanderhammeni brevipilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) tenuiscapa Westwood, 1840 (as Koptorthosoma tenuiscapa) 52c1cf8e550cbe7be9a31e9a128dc3bb taxon_sennertia_varicosa http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia varicosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Democratic Republic of the Congo: Katanga ([Monts] Kundelungu) (type locality), Sud-Kivu (Uvira); Angola "Minta-Luemba" 71399dcaa4fef654daf1212119378325 taxon_sennertia_varicosa http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia varicosa http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa (Mesotrichia) sp. (as Mesotrichia sp.) (type host) (Kundelungu); Xylocopa (Xenoxylocopa) inconstans Smith, 1874 (as Mesotrichia inconstans (Fain, 1971) and Mesotrichia flavescens inconstans (Fain, 1981a)) (Uvira and Minta-Luemba) 52facfe34a9eca45c04068114c4278bb taxon_sennertia_vitzthumi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia vitzthumi, phoretic deutonymph, ex Xylocopa latipes, Indonesia (Sumatra), BMOC 11-0930-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_vitzthumi_IRSNB_BMOC_11-0930-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 408fa946f206f787e808f3bbbe1793cd taxon_sennertia_vitzthumi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia vitzthumi, slide from IRSNB, BMOC 11-0930-002 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_vitzthumi_Slide_IRSNB_BMOC_11-0930-002.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e97c60d7c8d7ff901a896fe6218c58b6 taxon_sennertia_vitzthumi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia vitzthumi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Unknown (probably Oriental) (2 HDNs identified by Fain are from Inonesia (Sumatra), IRSNB (BMOC 11-0930-002)) ad354f7f4a92aee4ad410e1b93f5cea3 taxon_sennertia_vitzthumi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia vitzthumi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Unknown (2 HDNs identified by Fain are from Xylocopa latipes, IRSNB (BMOC 11-0930-002)) 4d9b9f921277755703b47627c35c666e taxon_sennertia_zhelochovtsevi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia zhelochovtsevi, lectotype, phoretic deutonymph, ex Xylocopa olivieri, Greece http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_zhelochovtsevi_lectotype.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a3c1b1cebbfc35420d8e84b2f674cbce taxon_sennertia_zhelochovtsevi http://purl.org/dc/dcmitype/StillImage image/jpeg Sennertia zhelochovtsevi lectotype slide http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Sennertia_zhelochovtsevi_lectotype_slide_BMOC_11-0618-003.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e da66aa7df2d047f26ae4792d8ba9f385 taxon_sennertia_zhelochovtsevi http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Sennertia zhelochovtsevi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Greece (type locality), Turkey, Armenia, "Middle Asian republics" of the former USSR (including Uzbekistan), Iran (our data, unpubl.). 1950b04cf0e4ae5724f52586d0227ed3 taxon_sennertia_zhelochovtsevi http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Sennertia zhelochovtsevi http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa olivieri Lepeletier, 1841 (type host); Xylocopa rufa Friese, 1901 d4da816be533c83a4836c361ad822df3 taxon_trigonholaspis http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis sp., ex Lestrimelitta limao, Panama, BMOC 88-0729-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_sp_BMOC_88-0729-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e a98e40f2371a569a6ad60d80b6290e2d taxon_trigonholaspis http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Trigonholaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description Setae J1, J2, J3 present; anterior dorsum with well-developed "hump" (Krantz, 1998). d2265fa29e3e3389789ba00d8b4b5cdf taxon_trigonholaspis http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Trigonholaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes four named species that inhabit nests of Trigona amalthea (Apidae: Meliponini) in Columbia (Vitzthum, 1930). Krantz (1998) reported five additional undescribed species from Brazil and Panama. We have seen an undescribed species from a nest of Lestrimelitta limao (Smith) from Panama. Individuals of this bee species rob nests of other bees, especially species of the genera Plebeia, Nannotrigona, more rarely Melipona, Scaptotrigona, and Trigona (Michener, 2000). The biology is almost unknown. Salt (1929) reported his observations onTrigonholaspis associated with Trigona amalthea (probably more than one species): "Mites are present on the comb in large numbers, as many as five or six together on the top of a single cell. In one or two cases I have seen these mites inside closed cells. On breaking open one cell in particular I found a bee pupa, its eyes only faintly colored, with a brown mite attached to it under the thorax apparently to the left middle leg; the mite was conspicuous as soon as the cell was opened. Mites are only rarely to be found on the pollen-and honey-pots, but on the comb are very numerous." 43a1b997c326e963a3b8e92fd1f56ae6 taxon_trigonholaspis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Genus Trigonholaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Neotropics. bfeddf2f55eb9b9075296ec5fa0a6ed0 taxon_trigonholaspis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Genus Trigonholaspis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Meliponine bees (Apidae: Meliponini). 14dbaa9d05e0488fa8a51c8653ae5ab8 taxon_trigonholaspis_amaltheae http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis amaltheae, female, ex Trigona amalthea, Colombia (after Vitzthum, 1930) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_amaltheae_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 03978419175e514a250e765dc73d7973 taxon_trigonholaspis_amaltheae http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis amaltheae, male, ex Trigona amalthea, Colombia (after Vitzthum, 1930) http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_amaltheae_m.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 434dbf1338c66f45284f7650803ce803 taxon_trigonholaspis_amaltheae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trigonholaspis amaltheae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Colombia: Magdalena (type locality) 10edb604fa3b4b85387c2be1dffa8dbf taxon_trigonholaspis_amaltheae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trigonholaspis amaltheae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona amalthea (Olivier, 1789). 54a447d89cbdb7433980525aeb1861d8 taxon_trigonholaspis_amaltheae http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Trigonholaspis amaltheae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected in the nest of Trigona amalthea. b75244011c365060af461b05f5bf8571 taxon_trigonholaspis_columbiana http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis columbiana, female, ex Trigona amalthea, Colombia (after Vitzthum, 1930) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_columbiana_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 335f30afa650af7c84a2e14126f15107 taxon_trigonholaspis_columbiana http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trigonholaspis columbiana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Colombia: Magdalena. b17c381a51c7c0ee4d91ae4304ebd259 taxon_trigonholaspis_columbiana http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trigonholaspis columbiana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona amalthea (Olivier, 1789). c0ab136da3d32cb6fca3f7e345358bd9 taxon_trigonholaspis_columbiana http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Trigonholaspis columbiana http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected in the nest of Trigona amalthea. 5db9e5007701ccfd648a1801771b0a3f taxon_trigonholaspis_salti http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis salti, female, ex Trigona amalthea, Colombia (after Vitzthum, 1930) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_salti_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e834beb9d66bfeb172ce48c1a80ab343 taxon_trigonholaspis_salti http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trigonholaspis salti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Colombia: Magdalena. a4563efc9b49a300f0275ce28d42bf2f taxon_trigonholaspis_salti http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trigonholaspis salti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona amalthea (Olivier, 1789). ac9a5e90c2aaf0adc5688e5526047144 taxon_trigonholaspis_salti http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Trigonholaspis salti http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected in the nest of Trigona amalthea. 228be4759abf7dee80a5beb7edff06d2 taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis sp., ex nest of the stingless bee Lestrimelitta limao, Panama, BMOC 88-0729-001 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_sp_BMOC_88-0729-001.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 929e89e054048287d3c6e81a30cd2c78 taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Panama: Colón. 5a3d1c2e1d6d0f1d3da3297db8ae7ba9 taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Lestrimelitta limao (Smith, 1863) (Apidae). e2df49532d2ff402b3281da834081f65 taxon_ http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected in the nest of Lestrimelitta limao. The mites were possibly originated from the nests of other stingless bees that have been robbed by L. limao. e47b9f573eeafcd367faf04f56272e26 taxon_trigonholaspis_trigonarum http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis trigonarum Vitzthum, 1930, female, ex nest of  the stingless bee Trigona amalthea, Colombia (after Vitzthum, 1930)  title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_trigonarum_f.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e d6141a6d12966863e6af65965a7d1f9a taxon_trigonholaspis_trigonarum http://purl.org/dc/dcmitype/StillImage image/jpeg Trigonholaspis trigonarum Vitzthum, 1930, male, ex nest of  the stingless bee Trigona amalthea, Colombia (after Vitzthum, 1930)  http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trigonholaspis_trigonarum_m.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 9bea2a3d0353a04a4ea2d06082df6d46 taxon_trigonholaspis_trigonarum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trigonholaspis trigonarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Colombia: Magdalena. d72f11e48e0695f94c68499b2cbb5893 taxon_trigonholaspis_trigonarum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trigonholaspis trigonarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Trigona amalthea (Olivier, 1789). 8e5618d63c79e64c110078657dd488b6 taxon_trigonholaspis_trigonarum http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Trigonholaspis trigonarum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Collected in the nest of Trigona amalthea. 00a37b7b86b92e16ced04e6326adb944 taxon_trochometridiidae http://purl.org/dc/dcmitype/StillImage image/jpeg Trochometridium tribulatum, phoretic female, ex Melissodes sp., USA: Arizona, BMOC 03-0821-006 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trochometridium_tribulatum_BMOC_03-0821-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2c957a78813d0e48a25de895151e5cb7 taxon_trochometridiidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Trochometridiidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This family includes two genera, Trochometridium Cross (Cross, 1965; Lindquist, 1986) and Neotrochometridium Hajiqanbar et Khaustov, 2009, associated with insects. Species of Trochometridium were collected from bees and bee nests (Hajiqanbar & Khaustov, 2009). 81873b3e31d0921fcbe2c5ecb74657f4 taxon_trochometridium http://purl.org/dc/dcmitype/StillImage image/jpeg Trochometridium tribulatum, phoretic female, ex Melissodes sp., USA: Arizona, BMOC 03-0821-006 http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trochometridium_tribulatum_BMOC_03-0821-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e e978d6bec6f4551717fa6c249318b66d taxon_trochometridium http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Trochometridium http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The genus includes five nominal species. Trochometridium tribulatum Cross is a polyxenous cleptoparasite of ground nesting bees in the Nearctic and Afrotropic regions. Trochometridium iranicum Hajiqanbar and Khaustov, 2009 was collected from the halictid bee Pseudapis nilotica (Smith, 1875) in Iran. Trochometridium chinensis (Mahunka, 1966) is a polyxenous cleptoparasite of wasps (China, New Guinea) and dermapterans (Iran). Trochometridium kazachstanicum Khaustov et Eidelberg, 2002 was found on carabid and scarabeid beetles in Kazakhstan and Iran, while Trochometridium kermanicum Mortazavi et Hajiqanbar, 2011 was described from an elaterid beetle in Iran. In addition, an undescribed species was recorded from Lasioglossum eremaean in Australia (Cross and Bohart, 1979; Hajiqanbar et al., 2009; Khaustov et Eidelberg, 2002; Lindquist, 1986; Mortazavi et al., 2009; Walter et al., 2002).
Biology is only known for Trochometridium tribulatum. 8a69308320b1c9115c5d2e5f78cc27ce taxon_trochometridium_iranicum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trochometridium iranicum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Iran (type locality - Iran, Razawi Khorassan Prov., vicinity of Sabzevar town, Mazinan village 36°18.510' N, 56°48.507' E, 877 m) 457da9faa86cadca69b74edd682fe549 taxon_trochometridium_iranicum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trochometridium iranicum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Pseudapis nilotica (Smith, 1875) 38610fb249b0cd391eda289639675e19 taxon_trochometridium_tribulatum http://purl.org/dc/dcmitype/StillImage image/jpeg Trochometridium tribulatum, phoretic female, ex Melissodes , USA: Arizona, BMOC 03-0821-006 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Trochometridium_tribulatum_BMOC_03-0821-006.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 2f268d3da3e70c5a59bb323e1e4cd11e taxon_trochometridium_tribulatum http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Trochometridium tribulatum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Canada: British Columbia; USA: Maryland, Idaho, California, Utah (type locality), Iowa, Kansas, Texas, Arizona, Alabama; Mexico: Tamaulipas; Egypt: Gizah; Sudan, South Africa: Gauteng (Cross and Bohart, 1979; Lindquist, 1985; our data). fa1b87a535996261c99bec3033b79fe6 taxon_trochometridium_tribulatum http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Trochometridium tribulatum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations (some records are probably accidental).
; North America:; Calliopsis (Nomadopsis) scutellaris Fowler, 1899; Calliopsis (Nomadopsis) anthidia Fowler, 1899; Calliopsis (Calliopsis) andreniformis Smith, 1853; Calliopsis (Calliopsima) coloradensis Cresson, 1878; Anthemurgus passiflorae Robertson, 1902 ; Halictus farinosus Smith, 1853 (type host); Halictus ligatus Say, 1837; Sphecodes arvensiformis Cockerell, 1904; Nomia (Acunomia) melanderi Cockerell, 1906; Nomia (Epinomia) nevadensis Cresson, 1874; Nomia nevadensis bakeri Cockerell, 1898; Megachile parallela Smith, 1853; Oreopasites scituli Cockerell, 1906; Exomalopsis birkmanni Cockerell, 1922; Melissodes fimbriata Cresson, 1878 (as "Melissoides fimbrioides"); Diadasia enavata (Cresson, 1872); Myrmosa unicolor Say 1824; Dasymutilla sp. (Hymenoptera: Mutillidae); Liris argentata (Beavois, 1811) (Hymenoptera: Sphecidae); Cicindela willistoni LeConte 1879 (Coleoptera: Carabidae). ; ; Afrotropic region: ; Aphodius sp. (Coleoptera: Scarabaeidae); Ossibia sp. (Coleoptera: Cerambycidae) ; ; (Cross and Bohart, 1979; Lindquist, 1985; Neff and Rozen, 1995; our data).;
eca798f79c53e3bdf9ab2b7eb3aaed31 taxon_trochometridium_tribulatum http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Trochometridium tribulatum http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology The host range includes andrenid, halictid, megachilid, and apid bees, as well as sphecid wasps and Coleoptera. Perhaps beetles become infested by mites dispersing from bees or wasps nesting in the same soil. The dispersing stage is the adult female. The female has a pair of internal sacs between legs III and IV that are termed sporothecae. Sporothecae contain fungal spores which are transferred by the mite to provisioned cells of ground nesting bees. So far the mite is known from the nests of bees of the genera of Calliopsis, Nomia, Halictus, and Anthemurgus. The bee egg or young larva dies in the infested cell as a result of development of the fungus and mites. Eggs are laid instead of being retained within the physogastric mother. These hatch into inactive larvae, which molt to males and females. The new generation of mites develops upon the fungal mycelium. The female is the only stage that feeds (Cross and Bohart, 1979; Lindquist, 1986; Neff and Rozen, 1995). 44f041d551cc8b7aea2700fc2984d25f taxon_tydeidae http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Family Tydeidae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This large cosmopolitan family includes mites that have been mostly recorded as predators but also as plant and fungus feeders or scavengers. Probably many species are unspecialized feeders. Tydeidae live in soil, humus, litter, moss, lichens, fungi, grass, on trees (in bark, on leaves and fruits), straw and hay, in stored products, and nests of birds, mammals, and stingless bees (Meliponini). Several species are of medical, veterinary, and agricultural importance. Tydeus molestus Moniez, 1889 has been recorded as causing itching and irritation of the skin of man and domestic animals.Tydeus interruptus Thor, 1932 was found in hay from Icelandic sheep pastures, and together with other mite species may represent a self-sustaining reservoir for scrapie-like agents. Tydeus californicus (Banks, 1904) and Lorryia formosa Cooreman, 1958 have been observed to cause damage to citrus. However, the last species is considered to be beneficial as it feeds on various fungi in citrus groves. (Kazmierski, 1998a; Krantz, 1978).
André (1980a, 1980b) listed 42 genera of the family, while Zhang at al., 2011 - 30 genera and 340 species. Kazmierski (1998a) gives a key to 21 genera and 254 species of the subfamily Tydeinae.
Many species of this family are known in associations with bees, although the exact nature of mite-bee interactions is unknown in these cases, but one species was speculated to be a beneficial fungus feeder in the host's nest. Two genera, Melissotydeus André and Proctotydaeus Berlese, were recorded from nests of stingless bees in Brazil. Unidentified species of Paralorryia and Pronematus were collected from beehives in Iran and the Philippines, respectively (Mosaddegh, 1997; Malabanan and Corpuz-Raros, 1988). Tydeus interruptus was found in bumblebee nests in New Zealand (Macfarlane, 2005), Tydeus xylocopae in the nest of a large carpenter bee in Indonesia (Oudemans, 1926), while unidentified species of Tydeus were repeatedly collected in beehives throughout the world (Chmielewski 1971, Haragsim et al. 1978, Homann 1933, Kamali et al. 2001, Malabanan and Corpuz-Raros 1998). Finally, a tydeid mite unassigned to a genus was recorded in beehives in Poland (Chmielewski, 1991; de Jong, 1982). 567b0ee9f5baddfee06be12a762c2615 taxon_tydeus http://purl.org/dc/dcmitype/Text text/html Description of the Bee Mite Genus Tydeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Description

Notes. There is no agreement over the year of the original description of Tydeus Koch. Most workers agree that the first description of Tydeus appeared in "Koch, C.L. Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur deutschen Fauna, Heft 4 (=Heft 137)." The most recent treatment cites the year as 1836 (André, 2005), with a discussion on publication date of Heft 4 - either 1836 or 1837 and the warrant that the year can not be found on the author's personal reprint. Oudemans (1937) gives the same year for Heft 4. However, the Nomenclator Zoologicus online database indicates that Heft 4 had been published in 1837. Kaźmierski (1989) gives 1835 for Tydeus Koch, without explicitly citing the original Koch's work or any further discussion on the matter. 1aa4b705eabb86588469e8f2d60a20f3 taxon_tydeus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Tydeus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription The majority of species of Tydeus are moss inhabitants (Momen and Lundqvist 2005). Some species occur on tree foliage, stored products, vertebrate nests and beehives. They are omnivorous scavengers and can also prey on the smallest arthropods and their eggs (Kaźmierski 2009). Unidentified species of Tydeus were repeatedly collected in beehives throughout the world (Chmielewski 1971, Haragsim et al. 1978, Homann 1933, Kamali et al. 2001, Malabanan and Corpuz-Raros 1998). Tydeus interruptus was found in bumblebee nests in New Zealand (Macfarlane, 2005), Tydeus xylocopae in the nest of a large carpenter bee in Indonesia (Oudemans, 1926).
The taxonomy of this genus is complicated, with many unrecognizable species described by earlier acarologists and the existence of several systematic treatments contradicting each other (André 2005). Major advances were made only recently, when re-examination of type species from Oudemans’ collection was made (André 2005). 1a46befd2c4b18f5fa1e19da459c20f2 taxon_tydeus_xylocopae http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Tydeus xylocopae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Indonesia: Medan: Deli c0d7f79ec5398e0a79ff675d0930eb1a taxon_tydeus_xylocopae http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Tydeus xylocopae http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Xylocopa sp. f69bc277f427723a6a7da73c5d5c4c9a taxon_urozercon http://purl.org/dc/dcmitype/StillImage image/jpeg Urozercon sp, female, ex nest of Scaura latitarsis, Costa Rica, BMOC 90-0101-020 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Urozercon_sp_BMOC_90-0101-020.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 32cb2f19f86df1d3c0bf05b645140635 taxon_urozercon http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Urozercon http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This little-known genus includes eight described species from Australia, South and Central America. Species of Urozercon occur commonly in tropical termite nests and have been observed riding on termites and stealing food during trophallaxis (Silvestri, 1917; Eickwort, 1990). Urozercon paradoxus Berlese, 1902, Urozercon cuneiformis (Trägårdth, 1906), Urozercon modestus Silvestri, 1917, Urozercon robustisetus (Rosario and Hunter, 1988), comb. n. (from Myrmozercon), Urozercon milleri (Halliday, 2006) comb. n. (from Uronyssus), and Urozercon watsoni (Halliday, 2006) comb. n. (from Uronyssus) were found in nests of termites. Several species were described from meliponine and centridine bees normally constructing their nests in termitaria: Urozercon melittophilus Silvestri, 1911 was found on the stingless bee Partamona cupira (Smith, 1863) while Urozercon angustatus Silvestri, 1911, stat. n. (Urozercon melittophilus var. angustatus Silvestri, 1911) was found on Centris thoracica (Hymenoptera: Apidae).
We collected a small series of Urozercon sp. in a nest of Scaura latitarsis (Friese, 1900) constructed inside a termite nest in Costa Rica (first record of Urozercon for North America). Silvestri (1903) reported nests of several species of Meliponini, including S. latitarsis, from nests of the termite Nasutitermes brevioculatus (Holmgren, 1910), the host of the mite Urozercon paradoxus. 37ae9ad570e3cff666572ab19d65c3a6 taxon_urozercon_angustatus http://purl.org/dc/dcmitype/StillImage image/jpeg Urozercon angustatus, ex Centris thoracica, Brazil (after Silvestri, 1911) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Urozercon_angustatus.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 00ec6edb193a756f9bd0cd1d2b29ae62 taxon_urozercon_angustatus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Urozercon angustatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Matto Grosso, Coxipó (type locality) 7e19d20701e54592a4e319aabe4b9436 taxon_urozercon_angustatus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Urozercon angustatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Centris (Melacentris) thoracica Lepeletier, 1841 (type host) d9e64ad6cddfa96bd31b47fa35a55238 taxon_urozercon_angustatus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Urozercon angustatus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found on the body of the host's larva. 837c6f35b77e8cf4676a318b85912730 taxon_urozercon_melittophilus http://purl.org/dc/dcmitype/StillImage image/jpeg Urozercon melittophilus, ex Partamona cupira, Brazil (after Silvestri, 1911) title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Urozercon_melittophilus.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e 4b04c6ff17fbf7623fd89309c4bb13f7 taxon_urozercon_melittophilus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Urozercon melittophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Brazil: Matto Grosso, Coxipó (type locality) 30c2022eb081478bbbb8858965c668cb taxon_urozercon_melittophilus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Urozercon melittophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Partamona cupira (Smith, 1863) (as Trigona cupira) (type host) d63e2d5eb832f606e849ac1ac1199f11 taxon_urozercon_melittophilus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Urozercon melittophilus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Found on the bee's metasoma. e2be6864b767a376e3881d0cfd531a3f taxon_ http://purl.org/dc/dcmitype/StillImage image/jpeg Urozercon sp, female, ex Scaura latitarsis, Costa Rica, BMOC 90-0101-020 title= http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Figs/Urozercon_sp_BMOC_90-0101-020.jpg http://creativecommons.org/licenses/by/3.0/ Bee-Associated Mite Database b7a9acd15655e25533d8b6974357860e d988eb1fc05b97ee9d590b19d2f19bcc taxon_ http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Costa Rica. 5761bef0226d97d396a7648a8860cbac taxon_ http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Scaura latitarsis (Friese, 1900) (Apidae: Meliponini). 6277a0107bbaccb5ba2b6e02acb24dc7 taxon_vulgarogamasus http://purl.org/dc/dcmitype/Text text/html General Description of the Bee Mite Genus Vulgarogamasus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription This genus includes 39 described species distributed in the New and Old World (Bai et al., 1995; Bai et al., 1991; El-Banhawy and Nasr, 1986; Hyatt, 1980; Gu and Guo, 1997; Gu and Huang, 1993; Gu et al., 1987; Karg, 1998; Ma, 1987; Ma, 1990; Ma et al., 2002; Ma-Liming and Wang-Shenron, 1996; Tichomirov, 1969a; Tseng, 1995; Ye and Ma, 1996; Ye et al., 1996). Many species of the genus are habitat generalists living in dung, decaying seaweed, forest litter, poultry litter, moss, and nests of small mammals and birds. One species, Vulgarogamasus sphecophilus (Cooreman, 1945), is restricted to the nests of wasps. Vulgarogamasus pollinerus El-Banhawy et Nasr, 1984 has been previously known only from a bee hive in Ithaca, New York. We found this species on bumblebees and in their nests. Vulgarogamasus cornutosimilis (Schweizer, 1949) (=Parasitus cornutosimilis (Karg, 1985; Micherdzinski, 1969)) is a little-known species described from Switzerland from Bombus terrestris and then recorded from Romania (forest litter) (Stănescu and Gwiazdowicz, 2004). Its two-pronged tectum resembles that of V. pollinerus. Finally, Vulgarogamasus aff. kraepelini was reported from honeybees in the Czech Republic (Haragsim et al., 1978) af03b3c8e93b2dfd58e392a468e2734e taxon_vulgarogamasus_cornutosimilis http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Vulgarogamasus cornutosimilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution Switzerland (type locality), Romania (Schweizer, 1949; Stănescu & Gwiazdowicz, 2004) 302cefd4cc874a9b33f89dbc30319dc6 taxon_vulgarogamasus_cornutosimilis http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Vulgarogamasus cornutosimilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations Bombus (Bombus) terrestris (Linnaeus, 1758) 0e2813c6dd1b0df6624ea06b92dfb548 taxon_vulgarogamasus_cornutosimilis http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Vulgarogamasus cornutosimilis http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology Probably a rare species. Found in moss, lichens, decaying wood, nest of Bombus (Bombus) terrestris in Switzerland (Schweizer, 1949), and forest litter in Romania (Stănescu & Gwiazdowicz, 2004). ecab8c84a82a0c87d676798325a03377 taxon_vulgarogamasus_pollinerus http://purl.org/dc/dcmitype/Text text/html Distribution of the Bee Mite Species Vulgarogamasus pollinerus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution USA; Indiana (first record); Michigan (first record); Montana (first record), New York (type locality). c1347ffa99c73af8e6a22b429e429ba2 taxon_vulgarogamasus_pollinerus http://purl.org/dc/dcmitype/Text text/html Host Associations of the Bee Mite Species Vulgarogamasus pollinerus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Associations
Apis mellifera Linnaeus, 1758 (type host); Bombus (Bombias) auricomus (Robertson, 1903) (first record); Bombus (Bombus) terricola Kirby, 1837 (first record); Bombus (Fervidobombus) pensylvanicus (De Geer, 1773) (as americanorum) (first record); Bombus (Pyrobombus) bimaculatus Cresson, 1863 (first record); Bombus (Pyrobombus) impatiens Cresson, 1863 (first record)
b272986973b20b4512e3039b17987d0b taxon_vulgarogamasus_pollinerus http://purl.org/dc/dcmitype/Text text/html Biology of the Bee Mite Species Vulgarogamasus pollinerus http://creativecommons.org/licenses/by-nc-sa/3.0/ b7a9acd15655e25533d8b6974357860e http://rs.tdwg.org/ontology/voc/SPMInfoItems#Biology A few deutonymphs were originally found in beehives in New York. We found seven deutonymphs phoretic on bumblebees and inside their nests. There is no evidence, however, that V. pollinerus is strictly associated with bees. Probably, this species, like other Parasitidae, preys on small arthropods in a variety of habitats, including bee nests.